PARISH CHECK LIST, LOUISIANA ODONATA

The Dragonflies And Damselflies (Odonata) Of Louisiana

by Bill Mauffray
International Odonata Research Institute
% Division of Plant Industry
P.0. Box 147100
Gainesville, FL. 32614
e-mail: iodonata@bellsouth.net

www.iodonata.net

Originally published in Bulletin of American Odonatology
Volume 5 Number 1
Oct 1997
Last update 27-June-2009
(updates since print publication in RED)

NEW FEATURE : Excellent photos and images mostly from Gayle and Jeanell Strickland of Baton Rouge to aid in field identifications.
Also Common names have been added (Sept 2004)

Abstract: The Dragonflies amd Damselflies (Odonata) of Louisiana

A comprehensive study of Louisiana dragonflies and damselflies (Odonata) was published by George Bick in 1957. He listed 101 species, including 33 Zygoptera and 68 Anisoptera. Since then several authors, including Mulhern (1971), Bick (1978 & 1990), Westfall & Tennessen (1979), and others (see Bibliography) have published data on an additional 14 species previously unreported in the state. This paper is an update and supplement to Bick's work.. It includes an update of the state list based on literature and specimens seen by the author. With 10 additional species and one removed from the list, there are now 124 taxa, including 33 Zygoptera, and 91 Anisoptera. Doubtful records and a list of species expected in Louisiana are also discussed. Quite a few distribution gaps have been filled within the last 40 years. A study of references from surrounding states has also helped to formulate a better understanding of the distribution of Odonata species within Louisiana. The bibliography includes Louisiana references, as well as that of surrounding states.

CONTENTS:

Introduction

Physiography Of Louisiana

Distribution Of Odonata Within Louisiana

Sources Of Information

Acknowledgments

The List (intro and legend)

Zygoptera

Anisoptera

Cordulegastridae

Corduliidae -Macromiinae

Corduliidae- Corduliinae

Libellulidae

Additions/Correstions To the List (since Jan 1998)

Doubtful Records

Expected Species

Bibliography

Introduction:

A comprehensive study of Louisiana dragonflies and damselflies (Odonata) was published by George Bick in 1957. He listed 101 species, including 33 Zygoptera and 68 Anisoptera. Since then several authors, including Mulhern (1971), Bick (1978 & 1990), Westfall & Tennessen (1979), and others (see Bibliography) have published data on an additional 14 species previously unreported in the state. This paper is an update and supplement to Bick's work. It includes an update of the state list based on literature and specimens seen by the author. With 10 additional species and one removed from the list, there are now 125 taxa, including 33 Zygoptera, and 92 Anisoptera. Doubtful records and a list of species expected in Louisiana are also discussed. Quite a few distribution gaps have been filled within the last 40 years. A study of references from surrounding states has also helped to formulate a better understanding of the distribution of Odonata species within Louisiana.

Although Louisiana has a variety of habitats for Odonata, It does not support the numbers of species that other southern and eastern states do: Alabama, 173 (Tennessen, et al,1995); Arkansas, 133 (Harp & Harp, 1996); Florida, 162 (Dunkle, 1992); New Jersey, 172 (May & Carle, 1996); New York 175 (Donnelly, 1992); Ohio, 156 (Glotzhober, 1995); and Texas, 194 (Abbot, in press). One primary reason for this could be that Louisiana is entirely below the fall line, and it's highest altitude is less then 600 feet above sea level.

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Physiography Of Louisiana:

All of Louisiana is in the Gulf Coastal Plain. Bick (1957) divided the state into 4 general physiographic zones: (see fig 1.) the Pinelands, Alluvial, Prairie, and Deltaic plain. This scheme seems to work out well with regards to Odonata distribution. The pinelands zone is separated by the alluvial areas of the Mississippi and Red rivers into 3 separate sub- areas in this study: (1) the Southeast Pinelands, extending eastward from the Mississippi alluvial area into southern Mississippi; (2) the Northern Pinelands, bounded on the east by the Mississippi alluvial area and on the west by the Red River alluvial area (this area extends northward into Arkansas and continues up to the Ouachita foothills); and (3) the Western Pinelands extends from the Red River alluvial areas westward to Houston and Tyler, Texas, and into southeastern Oklahoma.

INSERT FIG 1. PHYSIOGRAPHY OF LOUISIANA FILE NAME = LAPHYS2

INSERT FIG 2 PARISH MAP OF LOUISIANA FILE NAME= LAPARSH3

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Distribution Of Odonata Within Louisiana:

Bick (1957) gives an excellent account of distribution patterns within the state. Most of the Louisiana Odonata fauna is Eastern [E] (79= 64% ) or transcontinental [T] (19= 15. %). These 98 taxa (97 species, 2 subspecies) generally have a range extending from Florida and/or the Atlantic coastal states through Louisiana into east Texas or farther west. Twenty, or 16%, are Coastal Plain [CP], one is Western [W], and four have not been categorized [U]. No species is endemic to Louisiana, although one species, Ophiogomphus australis, is limited to the area that includes Washington and Tangipahoa Parishes, Louisiana, and Pike County immediately to the north in Mississippi.

Twelve species reach their western limits in Louisiana. Six of these species reach their western limit within the Washington-St.Tammany Parish area: Enallagma concisum, E. weewa, Gomphus hodgesi, Gomphus (Gomphurus) dilatatus, Cordulegaster bilineata, and Somatochlora provocans. Three taxa have been reported westward to the Mississippi alluvial area but do not cross it: Argia fumipennis fumipennis, Gomphus exilis, and Stylurus amnicola. Four species - Aphylla williamsoni, Dromogomphus armatus, Gomphaeschna antilope, and Libellula axilena- reach their western limit in south central and/or southwest Louisiana; however, there is an unverified record from southeast Texas for L. axilena (Orr, 1988).

Nine other eastern species have not been reported from western Louisiana but are expected there since they have been reported in southeast Texas: Hetaerina americana, Argia sedula, Enallagma daeckii, E. dubium, Gomphus apomyius, Anax longipes, and Neurocordulia alabamensis (Donnelly, 1978; Orr, 1988), and Epitheca spinosa from Arkansas and Oklahoma (Tennessen, 1994). Erpetogomphus designatus has also been reported only from this same area, but its range extends through Texas and also through Arkansas to the north. It would be expected in other parts of the state.

Four species reach their eastern limits in Louisiana: Gomphus militaris, Gomphus oklahomensis, Somatochlora margarita and Aphylla angustifolia. The latter is the only one of the four that has been found east of the Mississippi in Louisiana. The first three are limited to the pineland regions of the four state area consisting of Arkansas, Louisiana, Oklahoma, and east Texas. Two additional widespread taxa, Argia fumipennis violacea and Arigomphus submedianus, are reported from west of the Mississippi River but not from the southeastern part of the state, even though they are found on the east side of the river in the northern part of their range.

Only two species, Enallagma basidens and Epitheca stella, are restricted to extreme northern Louisiana. Libellula cyanea and L. luctuosa are primarily northern within the state, with a few recent records from the southern half of the state. Orr (pers. comm.) reports the latter from southeast Texas.

Thirty-six species were found primarily in the southern half or, at the most, the lower two thirds of the state. Erythrodiplax berenice and Macrodiplax balteata were found only in or near brackish areas. Miathyria marcella and Ischnura kellicotti were restricted by association with particular vegetation.

One species, Dythemis velox, which was not reported by Bick (1957), has apparently moved very rapidly from the north from Arkansas into north central Louisiana, and from the north through Mississippi into the northern part of the Florida Parishes of southeast Louisiana.

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Sources Of Information:

An attempt was made to review every published reference concerning Louisiana Odonata. The writer collected extensively in southeast Louisiana in the 60's and early 70's, and again from 1990-1992 in several regions of the state. Results of this field work are included here. The Jim Barr collection, LSU, IORI, FSCA, and USNM collections were personally studied. Data were provided from the personal notes of George & Juanda Bick, Sid Dunkle, Jerrell Daigle, John Abbott, and Ken Tennessen. In 1979, Jim Barr distributed a "list of Anisoptera" to the Louisiana Academy of Sciences. There were a number of discrepancies and some misdeterminations that were discovered when I studied his collection as well as the specimens at LSU. The results of this investigation are incorporated here. Malcolm Vidrine published a series of papers between 1988 and 1992 covering the Odonata of Louisiana, especially of the prairie region. Many of his records were based on sight records, and, for sake of proper citation, they are included, but voucher specimens are needed for verification.

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Acknowledgments:

George & Juanda Bick inventoried the FSCA collection and provided personal data, moral support, and the basis for this entire study. Nancy Adams assisted me in inventorying the USNM collection, Oliver Flint rechecked some pinned specimen determinations at the USNM, and Jim Barr provided his entire collection for study. Thanks also to John Abbott, Jerrell Daigle, Sid Dunkle, Geroge Harp, and Ken Tennessen for personal data. The staff of the LSU Entomology Collection, recently renamed the LSU Collection of Arthropods, allowed me to inventory their specimens. Dr. Minter J. Westfall Jr. provided additional moral support which helped rekindle my interest in Odonata after being inactive in Odonata studies for almost 20 years. George Bick, Jerrell Daigle, and Ken Tennessen reviewed the rough draft. Special thanks to Gayle and Jeanell Strickland for permission to link to their excellent images from the region. Credits for links to other images go to Dennis Paulson, and Giff Beaton.

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The List:

The families are listed after Garrison (1997) and all matters of nomenclature are based on his January 1997 revision. The species and parishes (= counties elsewhere) are listed alphabetically. All parish records included in and discussed by Bick (1957) are simply listed with no code. All parish records after 1957, plus any that were not covered by Bick (1957), are listed either with a code or a full citation. the latter is used if there were less than 3 records from the source. If a parish record not cited by Bick (1957) was from more than one source, then all the sources are listed.

In some cases a multiple-record reference may refer to the same specimen which was cited in the literature and then again in a particular collection. Many of the FSCA, IORI, USNM, and LSU records were collected by the writer. Some were in my personal collection at the beginning of this study but were later deposited in one of the studied collections. The multiple references better support the validity of the record, especially if one or more of the records was a sight record.

The earliest and latest dates for adults are enclosed in parentheses. Brackets that follow include codes for the general distribution of the species, restrictions within Louisiana, and a code for new species added since Bick (1957).

At the end of the above data are links to images, both male and female if available. Use the "back" button to return to this list.

Legend: codes within parentheses ( )

No code = All records (new and historical) from Bick (1957)

B= Bick personal communication

B8= Bick (1978)

B9= Bick (1990)

D= Dunkle (pers. comm)

G=Gayle & Jeanell Strickland 2003 and earlier

G4=Gayle & Jeanell Strickland 2004 records

G5=Gayle & Jeanell Strickland 2005 records

G6=Gayle & Jeanell Strickland 2006 records

G7=Gayle & Jeanell Strickland 2007 records

JL= Louton (1982)

F= FSCA/IORI collections (Gainesville, FL)

K4= Kreg Ellzey, per. comm (2004)

K5= Kreg Ellzey, per. comm.(2005)

K9= Kreg Ellzey, per. comm. (2009)

L= LSU (Baton Rouge, LA)

M= Mulhern (1971)

J= Barr Collection, verified by writer

JA= Abbott (pers. comm.)

JU= Barr 1979 List (unpublished)

O= R. Osborn (2005)

SS=Steve Shively 2007 records

S9=Steve Shively 2009 records

U= National Museum of Natural History (Washington DC)

V= Vidrine (1992)

V8= Vidrine (1988) from Ft. Polk list

V9= Vidrine (1991)

VP= Vidrine (1992) reported as "previous records" but unable to find a literature reference

W= Mauffray collection (many now in FSCA, IORI, LSU, and USNM)

WW= Walls & Walls (1971)

WT= Westfall & Tennessen (1979)

Legend: distribution codes within brackets [ ]

E= Eastern US

T= Transcontinental US

W= Western US

CP=Coastal Plain US

U= Undefined

> = Found only east of the Mississippi River in state

< = Found only west of the Mississippi River in state

^ = Found only in northern half of state

e = Reaches eastern limit in state

w = Reaches western limit in state

# = Added or reinstated to list since Bick (1957)

* = First record for state

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Zygoptera

Calopterygidae

Calopteryx dimidiata Burmeister, 1839. Sparkling Jewelwing. Allen (V), Beauregard, Rapides, St. Helena, St. Tammany, Tangipahoa (F,L), Union, Vernon (F,V,V8,W), Washington. Found along small, clear flowing streams in scattered localities throughout the pinelands. ( Mar 1-Sept 10) [E] Images: Male Female (Use <back> button to return to list )

Calopteryx maculata (Beauvois, 1805). Ebony Jewelwing. Allen (J,V), Beauregard, Bienville (J), Bossier, Caldwell (J), Catahoula (J), Claiborne, DeSoto (J), East Baton Rouge, East Feliciana, Grant (L,V), Jackson, LaSalle, Lincoln, Livingston (F,W), Morehouse (J), Natchitoches, Ouachita, Rapides, Red River (B), Sabine (B,J,V), St. Helena (F,J,W), St. Tammany, Tangipahoa, Union (J), Vernon (B,L,V8), Washington, West Feliciana (F,L,W), Winn. Very common throughout the pinelands along flowing creeks in mostly shaded situations.( Mar 1-Oct 25 ) [E] ] Images: Male Female (Use <back> button to return to list )

Hetaerina americana (Fabricius, 1798). American Rubyspot. East Baton Rouge, East Feliciana, St. Helena, Tangipahoa, Washington, West Feliciana (F,L,W). Reported only from the southeast pinelands along sunlit clear flowing streams. It should occur in other pineland areas, since it is reported from southeast Texas (Donnelly, 1978; Orr, 1988,1997). ( Apr 2-Oct 2 ) [T>] Images: Male Female (Use <back> button to return to list )

Hetaerina titia (Drury, 1773). Smoky Rubyspot. Allen, Avoyelles (VP), Beauregard (B,L), Catahoula (J), East Baton Rouge, East Feliciana, Evangeline, LaSalle (F,J), Madison, Natchitoches (L), Rapides, St. Helena, St. Tammany, Tangipahoa, Vernon, Washington, West Feliciana (F,L,W). This widespread species occurs throughout the pineland areas. ( Jun 24-Nov 15) [E] Images: Male Female (Use <back> button to return to list )

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Lestidae

Lestes australis Walker,1952. (=Lestes disjunctus australis ) Southern Spreadwing. Allen (V), Ascension, Beauregard (J), East Baton Rouge, East Feliciana (F), Iberville (L), Jefferson Davis (VP), LaSalle (F,W), Lincoln, Natchitoches (B), Ouachita, Rapides (B,V), Sabine, St. Landry(B,V), St. Tammany, Terrebonne, Washington (G5), West Feliciana (L). Collected in both pinelands and alluvial areas of the state, mostly around sunlit ponds, sloughs, and ditches. This species has two separate flight periods during the year. Bick (1957) suggested a dual emergence period. I found it rare in early spring but common in late fall. (Mar 8-Apr 4 and Aug 2-Dec 4) [E] Images: Male Female (Use <back> button to return to list )

Lestes Forficula Rambur, 1842. Rainpool Spreadwing. NEW STATE RECORD. St. Landry (G7). Images: Male Female (Use <back> button to return to list )

Lestes inaequalis Walsh, 1862. Elegant Spreadwing. Beauregard (V), East Feliciana (F,W), Grant (F), Natchitoches (K9), Saint Charles (F), Vernon (V), Webster. Except for the St. Charles records, collections were in the pinelands, but populations are apparently localized. (Apr 7-Aug 18) [E]

Lestes vigilax Hagen in Selys, 1862. Swamp Spreadwing. Acadia (V), Beauregard (V), Bossier, Caddo (L), Claiborne, East Feliciana (F), Jefferson Davis (VP), Lincoln, Rapides (V), St. Landry (V), St. Martin (V), St. Tammany, Tangipahoa, Union (J) Vernon (V). This shade-loving species was found in scattered locations only in the pinelands. (Mar 31-Oct 19) [E] Images: Male Female (Use <back> button to return to list )

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Coenagrionidae

Argia apicalis (Say, 1839). Blue-fronted Dancer. Acadia (V), Allen (J,V), Ascension, Avoyelles, Beauregard, Bienville, Caddo (K4), Calcasieu(D,F,L,V), Catahoula, Concordia (J), East Baton Rouge, East Carroll (J), East Feliciana, Evangeline (V), Grant, Jefferson Davis (J,V), Lafayette, LaSalle, Lincoln, Livingston (W), Madison, Morehouse (J), Natchitoches, Rapides (J,V), Sabine (V), St. Helena, St. Landry (V), St. Martin (V), Tangipahoa, Tensas, Washington, West Carroll (J), West Feliciana. Bick (1957) stated that this species was found almost equally in the pinelands and alluvial areas. (Apr 17-Nov 24) [E] Images: Male Female Female [brown form] (Use <back> button to return to list )

Argia bipunctulata (Hagen, 1861). Seepage Dancer. Caldwell, DeSoto, Lincoln, Livingston (W), Vernon (W), Washington (J). This widespread species is usually found in association with seepage springs which are not too abundant in Louisiana. Previous records were only for North Louisiana, but this species has now been collected in the pinelands of both the southeast and southwest. The Vernon Parish record was listed as Argia barretti by Vidrine et al (1992). See notes under "Doubtful and Obsolete Records" below. (May 5-Aug 4) [E] Images: Male Female (Use <back> button to return to list )

Argia fumipennis fumipennis (Burmeister, 1839). Smoky-winged Dancer. East Baton Rouge, East Feliciana, St. Helena, St. Tammany, Tangipahoa, Washington , West Feliciana (W). Found only in the southeast pinelands in a variety of habitats, sometimes being very abundant. (Mar 28-Oct 4) [E>] Images: Male Female (Use <back> button to return to list )

Argia f. violacea (Hagen, 1861). Violet Dancer. Bienville (J), Claiborne, LaSalle (F,W), Natchitoches (J), Rapides (B,V,W), Vernon. This clear winged form is found only west of the Mississippi in Louisiana, never as abundant as its southeast Louisiana dark winged form. The Bick(1957) records and those of the writer were from ponds. My experience with this species in Arkansas and Arizona is that it is a stream species. ( Mar 26-Sep 1 ) [E<] Images: Male (Use <back> button to return to list )

Argia moesta (Hagen, 1861). Powdered Dancer. Acadia (VP), Allen, Assumption (VP), Avoyelles (V), Beauregard, Bienville (JA), Claiborne, East Baton Rouge, East Feliciana, Evangeline (V), Grant, Jefferson Davis (J), LaSalle, Livingston, Madison (VP), Morehouse, Natchitoches (J), Rapides, Sabine (V), St. Helena, St. Tammany, Tangipahoa, Tensas (VP), Union (J), Vernon, Washington, West Feliciana (W). Winn. Found on streams and rivers throughout the pinelands. ( Apr-3-Sep 11) [T] Images: Male , Male (color var ) , Female , Female (color var.) (Use <back> button to return to list )

Argia sedula (Hagen, 1861). Blue-ringed Dancer. Caddo (O) ,East Baton Rouge, East Feliciana, Grant (B), LaSalle, Livingston, Madison, Richland (J), St. Helena, St. Tammany, Tangipahoa, Tensas, Washington, West Feliciana. This stream species is widespread but not as common as Argia moesta. It has not been reported from western Louisiana, but it should occur there. ( Apr 17-Sep 25 ) [T] Images: Male Female (Use <back> button to return to list )

Argia tibialis (Rambur, 1842). Blue-tipped Dancer. Acadia (L,V), Allen, Ascension, Avoyelles (V), Beauregard, Bienville (J), Bossier, Caddo, Calcasieu (V), Caldwell (J), Catahoula, Claiborne, DeSoto, East Baton Rouge, East Carroll (J), East Feliciana, Evangeline, Grant, Jackson, Jefferson Davis, Lafayette (B,V), LaSalle, Lincoln, Livingston, Madison, Morehouse, Natchitoches, Rapides, Red River (J), Sabine (V), St. Helena, St. Landry, St. Martin (V), St. Tammany, Tangipahoa, Union, Vernon, Washington, Webster, West Carroll, West Feliciana (W), Winn. Very widespread on both pond and stream habitats, but absent from the Deltaic Plain. (Mar 3-Sep 16) [E] Images: Male Female (Use <back> button to return to list )

Enallagma basidens Calvert, 1902. Double-striped Bluet. Caddo (O), East Baton Rouge (G), Natchitoches, Tensas, West Feliciana (G5). So far, basidens has been found only in northern Louisiana. This species has rapidly expanded its range from southwestern US into the mid-west and north Atlantic states. It should be found at other localities in northern and western Louisiana. ( Apr 04 -Oct 5) [T^] Images: Male Female (Use <back> button to return to list )

Enallagma civile (Hagen, 1861). Familiar Bluet. Acadia (V), Allen (V), Ascension (W), Beauregard (V), Caddo (O), Calcasieu (V), Caldwell, Cameron (G), Catahoula (J), Concordia (J), East Baton Rouge, East Feliciana (J), Evangeline (B), Iberia (B), Jackson, Jefferson, Jefferson Davis (V), LaSalle, Lincoln, Livingston (W), Madison, Natchitoches, Orleans (F,U), Ouachita, Sabine (V), St. Bernard (W), St. Helena (W), St. Landry (V), St. Tammany (F), Tangipahoa (W), Tensas, Vernon (V), Webster, West Feliciana. This species was found in a variety of habitats throughout the state including slightly brackish ponds and flowing streams. (Mar 8-Nov 26 ) [T] Images: Male Female (Use <back> button to return to list )

Enallagma concisum Williamson, 1922. Cherry Bluet. St. Tammany, Jefferson. The latter record is from a brackish marsh at Grand Isle, Louisiana. This is unusual, since E. concisum is usually associated with heavily vegetated, acidic ponds and roadside ditches in the pinelands of southeastern US. The Grand Isle specimen could have been a vagrant blown in from Saint Tammany Parish, where it has been found only in the eastern half of the parish. These are the westernmost records for the species. ( Mar 6-Sep 27) [CP>w] Images: Male Female (Use <back> button to return to list )

Enallagma daeckii (Calvert, 1903). Attenuated Bluet. St. Tammany, Tangipahoa (W). Found only in a few scattered localities of the southeast pinelands. (Apr 24-Jun 8) [CP>] Images: Male Female (Use <back> button to return to list )

Enallagma divagans Selys, 1876. Turquoise Bluet. Bienville (J), Catahoula (J), East Baton Rouge (W), East Feliciana (W), Livingston (F), Natchitoches, Rapides (B), St. Helena, St.Tammany, Tangipahoa, Vernon (V,V8), Washington (D,W). Common early season species found throughout the pinelands. ( Mar 14-Jun 12) [E] Images: Male Female (Use <back> button to return to list )

Enallagma doubledayi (Selys. 1850). Atlantic Bluet. NEW STATE RECORD: Natchitoches (G4, K5), St. Tammany (G4) (Apr 8–Jun 13) [E] Images: Male Female (Use <back> button to return to list )

Enallagma dubium Root, 1924. Burgundy Bluet. Acadia (V), Calcasieu (V), Jefferson Davis (VP), St. Tammany. Although the Vidrine (1992) records are sight records, E. dubium probably does occur in southwest Louisiana, as Donnelly (1978) and Orr (1988) reported it from southeast Texas. (Apr 9-Sep 10) [CP] Images: Male Female (Use <back> button to return to list )

Enallagma durum (Hagen, 1861). Big Bluet. Calcasieu (JA), Cameron (L,V), East Baton Rouge (W), Jefferson, Lafourche (F), Madison, Orleans, Plaquemines (B), St. Charles (W), St. Tammany (F,L). This species was found at various sites within the deltaic and alluvial zones. The St. Tammany site at Slidell is on the edge of the pinelands. Some sites are slightly brackish. ( Apr 1-Oct 25) [CP] Images: Male Female (Use <back> button to return to list )

Enallagma exsulans (Hagen, 1861). Stream Bluet. Allen (B), East Feliciana (W), Jefferson Davis (V), LaSalle (W), Madison, Natchitoches (B), St. Helena (F), St. Landry (V), Tangipahoa (G5) ,Tensas (J), Washington . Found along clear flowing streams in the pinelands. (Apr 12-Sep 10) [E] Images: Male Female (Use <back> button to return to list )

Enallagma geminatum Kellicott, 1895. Skimming Bluet. Evangeline (B), Madison, St. Tammany (L,W), Tensas, Webster, West Feliciana (W). This widespread species apparently occurs only locally in Louisiana. ( Mar 31-Aug 18) [E] Images: Male Female (Use <back> button to return to list )

Enallagma signatum (Hagen, 1861). Orange Bluet. Acadia (V), Ascension, Avoyelles (J), Beauregard (B), Caddo (J, O), Calcasieu (D,V), Caldwell, Cameron, Catahoula, Claiborne, East Baton Rouge, East Feliciana (W), Evangeline, Grant (W), Iberia (B), Iberville (W), Jefferson, Jefferson Davis (V), Lafourche, LaSalle (W), Lincoln, Livingston (W), Madison, Natchitoches, Orleans, Pointe Coupee (F), Rapides, St. Helena (F,W), St. Landry (V), St. Martin, St. Tammany, Tangipahoa, Tensas, Vernon (V,V8), Webster, West Baton Rouge (W), West Carroll (J), West Feliciana. This species occurred in all physiographic regions. (Feb 13-Nov 10) [E] Images: Male Female (Use <back> button to return to list )

Enallagma traviatum westfalli Donnelly, 1964. Slender Bluet. Tangipahoa, Vernon (V,W) Washington (D), West Feliciana (G5). Reported as E. traviatum by Bick (1957). This is another species that has very localized populations in the state. The one in Tangipahoa is an old gravel pit adjacent to the Tangipahoa River at Amite. Sid Dunkle reports (pers. comm.) that the Washington Parish specimens (2 males) were collected in the forest, not at the water. The locality was along the Pushepatapa Creek at SR-21 near Varnado.( Apr 17-May 3) [E] Images: Male1 Male2 Female (Use <back> button to return to list )

Enallagma vesperum Calvert, 1919. Vesper Bluet. Beauregard (G5), Cameron (G5), Claiborne, St. Tammany (W), Tensas, West Feliciana (G5). The St. Tammany site is a wooded, marshy edge of a spring-fed man-made pond at Florenville. ( Mar 31-Aug 17) [E] Images: Male Female (Use <back> button to return to list )

Enallagma weewa Byers, 1927. Blackwater Bluet. St. Tammany. This is the westernmost record for this species, because the Allen (V), Rapides (V), and Vernon (V8) records are questionable. It is unlikely that this species occurs west of the Mississippi River. It is usually found hovering almost on the surface of deeply stained acid-water pools of flowing streams, where the adults are difficult to see and difficult to capture unless "pancaked" with a net. ( Apr 25-Sep 10) [CP>w]

Ischnura hastata (Say, 1839). (= Anomalagrion hastatum). Citrine Forktail. Acadia (B,J,V), Allen, Ascension (W), Avoyelles(U), Beauregard (B,J,V), Caddo (O), Calcasieu (U,V), Caldwell, Cameron, Catahoula (J), Claiborne, East Baton Rouge, East Carroll (J), East Feliciana (W), Evangeline (V,W), Franklin, Grant, Iberia, Jackson, Jefferson, Jefferson Davis (V), Iberville (L), Lafayette, LaSalle, Lincoln, Livingston (W), Madison, Morehouse, Natchitoches, Orleans, Ouachita, Plaquemines, Rapides, Richland, Sabine, St. Charles (B). St. John the Baptist (L), St. Helena, St. James (V,W), St. Landry (L), St. Martin, St. Tammany, Tangipahoa, Tensas, Union, Vermilion, Vernon, Washington, Webster, West Baton Rouge (W), West Feliciana (J), Winn. ( Feb 17-Nov 18) [T] Images: Male Female, Young Female (Use <back> button to return to list )

Ischnura kellicotti Williamson, 1898. Lilypad Forktail. Caddo (O), Claiborne, East Feliciana (L), Lincoln, Ouachita, St. Tammany, Vermilion (J). Should be found in the pinelands wherever there are lentic, acid waters with Nuphar or Nymphaea pads. Bick (1957) discussed the method of scooping up the whole lily pad to collect the adults. Ken Tennessen recommends (pers. comm.) simply placing the net slowly over the lily pad and allowing the specimen to fly up into the net. ( Apr 1-Oct 1) [E] Images: Male Female (Use <back> button to return to list )

Ischnura posita (Hagen, 1861). Fragile Forktail. Acadia, Allen (B,V), Ascension, Avoyelles (J), Beauregard (B,V), Bienville (J), Caddo (J, O), Calcasieu (U,V), Caldwell, Cameron (V), Catahoula, Claiborne, Concordia (J), East Baton Rouge, East Carroll (J), East Feliciana, Evangeline, Franklin, Grant, Iberia (B,V), Iberville (W), Jackson, Jefferson, Jefferson Davis (V), Lafayette, Lafourche, LaSalle, Lincoln, Livingston, Madison, Morehouse, Natchitoches, Orleans, Ouachita, Plaquemines, Point Coupe (W), Rapides, Richland, Sabine, St. Charles, St. Helena (W), St. John the Baptist (F,L), St. Landry (V), St. Martin, St. Mary (J), St. Tammany, Tangipahoa, Tensas, Terrebonne, Vermilion (V), Vernon, Washington, Webster, West Baton Rouge (W), West Feliciana, Winn (B). ( Feb 13-Dec 31) [E] Images: Male Female (Use <back> button to return to list )

Ischnura prognata (Hagen, 1861). Furtive Forktail. Cameron, East Baton Rouge (W), Jefferson, Orleans, St. John the Baptist (F), St. Martin (B), St. Tammany, Tangipahoa. This shade-loving species was found in isolated swampy habitats across southern Louisiana. ( Feb 15-Sep 9) [E]

Ischnura ramburii (Selys, 1850). Rambur's Forktail. Acadia (D,L,V), Allen, Ascension, Avoyelles, Beauregard, Bienville, Bossier, Caddo (O), Calcasieu, Cameron, Claiborne, East Baton Rouge, East Carroll, East Feliciana, Evangeline (B,V), Franklin, Iberia, Iberville (L), Jackson, Jefferson, Jefferson Davis, Lafayette, Lafourche, LaSalle (W), Lincoln, Livingston, Madison, Natchitoches, Orleans, Ouachita, Plaquemines, Point Coupe (J), Rapides, Richland (J), St. Bernard (W), St. Helena (W), St. James (W), St. John the Baptist (F), St. Landry (V), St. Martin (B,V), St. Mary, St. Tammany, Tangipahoa, Tensas, Terrebonne, Vermilion, Vernon (V,V8), Washington (W), West Baton Rouge (W), West Feliciana. It occurs throughout the state but is more common in the southern third. ( Jan 2-Dec 23) [E] Images: Male Female Female red form (Use <back> button to return to list )

Nehalennia integricollis Calvert, 1913. Southern Sprite. Natchitoches (K4), St. Tammany, Calcasieu (L). Sam Houston State Park 17-V-1992, 1 female, coll. J. Louton. These records are at the far end of south Louisiana. Additional collecting should result in more localities. ( Mar 31-Sep-10) [E] Images: Male Female (Use <back> button to return to list )

Telebasis byersi Westfall, 1957. Duckweed Firetail. Lousiana (Westfall & May, 1996). Iberville (G5), St. John the Baptist: This record is based on a young (3-4th instar) larva in the FSCA collection with the following data" "Reserve Canal","26-IX-73" No collector was listed. This locality in St. John the Baptist Parish is a muddy drainage canal in the midst of sugarcane fields. The specimen was originally determined as T. salva, but when it was examined again for this study in 1996, It was redetermined as byersi. When I checked the Reserve Canal site in the mid 70's, it was devoid of vegetation and not accessible for any larval sampling. Vidrine et al, 1992 reports sight records from Beauregard, Lafayette and St. Landry Parishes. It has been recorded from East Texas (Orr, 1988; Dunkle, pers. comm.) and southern Arkansas, Drew Co (Harp & Harp, 1996), so it probably does occur in other parts of Louisiana. Voucher specimens are needed. [#E] Images: Male Female (Use <back> button to return to list )

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Anisoptera

Petaluridae

Tachopteryx thoreyi (Hagen in Selys, 1858). Gray Petaltail. Grant (F), Natchitoches (J), Rapides (V), Sabine (V), St. Tammany, Union, Vernon (D,V,V8,V8b). This widespread eastern species is rare in Louisiana, due to the scarcity of seepage springs. The Grant parish specimen was collected as it landed on my white T-shirt. ( Apr 3-Jul 15) [E] Images: Male Female (Use <back> button to return to list )

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Aeshnidae

Anax junius (Drury, 1770). Common Green Darner. Acadia (V), Allen, Ascension (JU), Avoyelles (M,V), Beauregard, Bossier (L,JU), Caddo (L,JU,O), Calcasieu, Cameron, Catahoula (J), Claiborne, Concordia, DeSoto (L,JU), East Baton Rouge, East Feliciana (F,JU), Evangeline, Franklin, Iberia, Iberville, Jefferson, Jefferson Davis (L,JU), Lafayette, LaSalle (W), Lafourche, Lincoln, Livingston (JU,W), Madison, Natchitoches, Orleans, Ouachita, Pointe Coupee (JU), Plaquemines, Rapides, Red River (JU), Richland, Sabine (V), St. Bernard, St. Charles, St. Helena (F,W), St. John the Baptist (F,L), St. Landry (L,JU), St. Martin, St. Mary, St. Tammany, Tangipahoa, Terrebonne, Vermilion, Vernon, Washington, Webster (L,JU), West Carroll (J), West Baton Rouge (JU), West Feliciana. Bick (1957) reported a swarm of about 1000 adults over the marshes of Grand Isle in the Gulf of Mexico. ( Jan 19-Dec 14) [T] Images: Male Female (Use <back> button to return to list )

Anax longipes Hagen, 1861. Comet Darner. Natchitoches (K4), Ouachita (M), St. Tammany, Tangipahoa (B), West Feliciana (V). ( May 3-Aug 30) [E] Images: Male (Use <back> button to return to list )

Basiaeschna janata (Say, 1839). Springtime Darner. DeSoto (JU), Livingston (JU), Ouachita (M), Rapides (B,D), St. Helena (F,L,W), Tangipahoa (F), Union (JU), Washington, West Feliciana (JU). This early season species was found along small to medium clear flowing streams in widely scattered locations throughout the pinelands ( Mar 26-Apr 6) [E] Images: Male Female (Use <back> button to return to list )

Boyeria vinosa (Say, 1839). Fawn Darner. Allen (JU,WW), Beauregard (JL,V), Bossier, Caldwell (J), DeSoto (VP), East Feliciana, Evangeline (V), Grant, Jefferson Davis (JL), LaSalle (J), Livingston, Natchitoches, Rapides, St. Helena, St. Tammany, Tangipahoa, Union (J,L), Vernon, Washington, West Feliciana. Bick (1957), May & Carle (1996) considered the scarcity of adults due to their crepuscular habit. I collected adults while wading clear flowing streams along deeply shaded water courses and found the adults erratically flying along the shoreline. (May 3-Nov 5) [E] Images: Male Female (Use <back> button to return to list )

Coryphaeschna ingens (Rambur, 1842). Regal Darner. Acadia (D,V), Allen, Beauregard (V), Caddo (K4), Calcasieu (D), Cameron, Claiborne (U), DeSoto (U), East Baton Rouge, East Feliciana (L,JU), Evangeline, Jefferson, Jefferson Davis (D), Lafayette, Morehouse (JU), Natchitoches (K4), Orleans, Plaquemines, Rapides (S9), St. Bernard, St. Charles, St. Helena, St. James, St. John the Baptist (F), St. Landry (V), St. Mary (J,JU), St. Tammany, Tangipahoa, Terrebonne, Vermilion (VP), Vernon.. Most records are from the southern half of the state in pinelands, alluvial, and deltaic areas. I observed swarms of this species in Jefferson Parish at dusk on several occasions during August of 1965. ( Apr 15-Oct 3) [E] Images: Male (Use <back> button to return to list )

Epiaeschna heros (Fabricius, 1798). Swamp Darner. Acadia (D,JU), Allen, Ascension (W), Assumption (VP), Avoyelles (VP). Beauregard (VP), Bienville (JU), Bossier, Caddo, Calcasieu (VP), Caldwell (VP), Cameron, Catahoula (VP), Claiborne (VP), Concordia (L), DeSoto (VP), East Baton Rouge, East Carroll (VP), East Feliciana, Evangeline, Franklin (J), Grant, Iberville, Jackson, Jefferson, Jefferson Davis (D,VP), Lafayette, Lafourche, LaSalle (W), Lincoln (VP), Livingston, Madison (VP), Morehouse (J), Natchitoches, Orleans, Ouachita (VP), Point Coupe (L, J), Rapides (B,D), Red River (VP), Sabine (V), St. Bernard (F), St. Charles, St. Helena (JU,W), St. James (VP), St. John the Baptist (D,F), St. Landry, St. Martin (JP), St. Mary (VP), St. Tammany, Tangipahoa, Tensas (VP), Terrebonne, Union (J), Vermilion (B), Vernon (B,D,V8), Washington, Webster, West Baton Rouge (JU), West Carroll (VP), West Feliciana, Winn. There are many discrepancies in the "Previous Records" reported by Vidrine (1992) [as "VP"] as compared to actual literature records. I compared the Vidrine "Previous Records" with the Jim Barr unpublished handout (1979) and found exactly the same discrepancies. Perhaps Barr somehow confused the "previous records" for this species with those listed for another species. Vidrine simply repeated the Barr List. It probably will eventually be found in all these additional parishes as it was not restricted to any zone. ( Feb 22-Oct 28) [E] Images: Male Female (Use <back> button to return to list )

Gomphaeschna antilope (Hagen, 1874). Taper-tailed Darner. Lafayette (B), Orleans, St. Tammany (D,F). Both species of this genus were found near swampy areas. The Lafayette Parish record is the westernmost for this species. ( Apr 11-Jun 2) [Ew]

Gomphaeschna furcillata (Say, 1839). Harlequin Darner. Jackson, Lafayette (F), Madison, Natchitoches (G6), Orleans, Rapides (D). ( Feb 3-Apr 14) [E] Images: Male Female (Use <back> button to return to list )

Nasiaeschna pentacantha (Rambur, 1842). Cyrano Darner. Acadia (D,V), Allen (W), Ascension, Assumption, Caddo (JU,O), DeSoto, East Baton Rouge, East Feliciana (G5), Evangeline (V), Iberia (V), Iberville, Jefferson, Jefferson Davis (D), Lafayette,Lafourche (JU), Madison, Natchitoches, Orleans, Plaquemines, Rapides (V), Richland, Sabine (V), St. Charles, St. Landry (V), St. Martin (V), St. Tammany, Tangipahoa, Washington, West Baton Rouge (JA), West Feliciana (W). (Mar 30-Oct 12) [E] Images: Male (Use <back> button to return to list )

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Gomphidae

Aphylla angustifolia Garrison,1986. Broad-striped Forceptail. West Feliciana: Laurel Hill; Lake Rosemond development, "N.E. corner, near small lake": 22-July-1990. 1 male (W); East Baton Rouge: Central, Biltmore Subdivision: 22-June-1986,1 female (J) same: 2-July-1984: 1 male (J); Allen Kinder, Pond, 4-July-1968, M. Milson "Data from Jerry Walls". (per. comm. Sid Dunkle), Rapides (S9). Vidrine, (1988a,1992) reported sight records of A. protracta (Selys, 1859).from Calcasieu, Evangeline, Jefferson Davis, Lafayette, St. Landry and Vermilion. These are probably angustifolia. A. angustifolia apparently has become distributed in Louisiana from west to east into the Baton Rouge and West Feliciana areas. The West Feliciana site at Lake Rosemond is a man-made sand-bottomed lake which also supports an A. williamsoni population. ( Jun 9- Aug 14) [*We] Images: Male (Use <back> button to return to list )

Aphylla williamsoni (Gloyd, 1936). Two-striped Forceptail. Beauregard, East Baton Rouge, Iberia (V), East Feliciana (G5), Jefferson, Lafourche (JU), Natchotoches (K4), Orleans, Sabine (V), St. Bernard (F), St. Charles, St. Landry (V), St. Mary (JU), St. Tammany, West Feliciana (W). This species is one of the few gomphids found in the deltaic plain and the only one in the New Orleans area. The Beauregard and Sabine records are the westernmost for this species. ( Apr 14-Nov 2) [Ew] Images: Male Female (Use <back> button to return to list )

Arigomphus maxwelli (Ferguson, 1950). Bayou Clubtail. Acadia: ditch, 1 ml E of Jefferson Davis Parish line on I-10, 15-June-1966, 1 male, (D,F); Calcasieu: pond, 6.2 ml E. Vinton turnoff of I-10, 15-June-1966, 1 male (D,F); Claiborne: 17-June-1979. 1 male (J); East Baton Rouge: Klienpeter, pond, 1 ml W of US-61 & LA 42. 24-May-1967, 1 male; ibid 17-June-1966, 1 male (F,W); Jefferson Davis: Bayou Nezpique at Hwy 97, 2-May-1985, 1 male(D,F); Lafayette (B); St. Martin: East edge of Henderson at Bayou Amy, 2 June 1997, 8 males, 1 female (George Harp, pers. comm.); St. Landry (G7), Tangipahoa: 5-June-1977, 1 male (L). These seem to be the first verified Louisiana records. ( May 2-Jun 17) [*U] Images: Male Female (Use <back> button to return to list )

Arigomphus submedianus (Williamson, 1914). Jade Clubtail. Acadia (J), Caddo (L), Morehouse, Natchitoches (G), Point Coupee (J), Rapides, Tensas (J). (May 23-Jul 31) [E] Images: Male Female (Use <back> button to return to list )

Dromogomphus armatus Selys, 1854. Southeastern Spinyleg. Iberia (WT), St. Tammany (JL). Only records are larvae. The Iberia record is the westernmost for this species. The St. Tammany record was from Talisheek Creek @ LA-41. [#CPw] Images: Male (Use <back> button to return to list )

Dromogomphus spinosus Selys, 1854. Black-shouldered Spinyleg. Acadia (JU), Allen, Avoyelles (V), Beauregard (JL,V), Bienville (JA), Bossier (JU), Caddo (JU,L), Caldwell (J), Catahoula (J), Claiborne (JU,L), DeSoto (JU), East Baton Rouge (Garcia, 1934,JU,L), East Feliciana, Evangeline (V), Grant, Iberia, Jackson (JU), Jefferson Davis (JL,JU), LaSalle (JL), Livingston, Madison, Morehouse (VP), Natchitoches (K9), Ouachita (M), Pointe Coupee (JU), Rapides, Sabine (JU), St. Helena, St. Landry (V), St. Martin (V), St. Tammany, Tangipahoa (G5) ,Union (J,JU), Vernon (JU,L), Washington (JL), West Feliciana (JU). I have redetermined the Webster specimen (JU), in the LSU collection, as D. spoliatus, and included it under that species. (May 24--Sep 11) [E] Images: Female Male (Use <back> button to return to list )

Dromogomphus spoliatus (Hagen in Selys, 1858). Flag-tailed Spinyleg. Acadia (V), Allen (D,WT), Concordia (G), Evangeline (V), Jefferson Davis (V), Morehouse (M), Rapides (V), Sabine (JL). St. Landry (V), Vernon (V). Webster (L). Bick(1957) reported larvae from 6 additional parishes, but he stated (pers. comm.) that the identifications may not be valid. ( Jun 19-Aug 12) [E] Images: Male (Use <back> button to return to list )

Erpetogomphus designatus Hagen in Selys, 1858. Eastern Ringtail. Caddo (O), St. Tammany (F), Washington. This widespread species has only been collected in the Pearl River drainage. Further collecting should result in additional records. E. designatus is widespread throughout Texas, Oklahoma, Arkansas, and Mississippi. The St. Tammany specimens were collected at the town of Pearl River, near the West Pearl River bridge at US-11 and I-10 on 6-June-1964. [E>] Images: Male Female (Use <back> button to return to list )

Gomphus (Gomphurus) dilatatus Rambur, 1842. Blackwater Clubtail. Foster & Smith (1901) originally recorded this species "from the Amite River." Bick (1957) listed it in his "species not seen by the writer" category and discussed the historical notes about the specimen. Westfall (1974) determined the Amite River specimen as G. modestus. G. dilatatus has been collected in Washington Parish: Crain Creek, 28-May-1987,1 female, coll Jim Barr. This specimen has been checked by me. Another record is from Varnado, Pushepatapa Creek, SR 21, 3-May-1986, 2 males coll. by Sid Dunkle. G. dilatatus seems to reach its western boundary in the southeast pinelands.[*CP>w] Images: Male Female (Use <back> button to return to list )

Gomphus (Gomphurus) hybridus Williamson,1902. Cocoa Clubtail. Allen (W), Beauregard (JL), DeSoto(U), East Baton Rouge (W), East Feliciana (W), St. Tammamy (F), Tangipahoa(D) Washington (D). Sight records from Vidrine for Gomphus vastus are probably this species. The writer spent a day collecting with him in March, 1992 at his vastus location, and we got only this species. ( Mar 30-May 3) [#E] Images: Male Female (Use <back> button to return to list )

Gomphus (Gomphurus) modestus Needham,1942. Gulf Coast Clubtail. Allen (WW), Rapides (S9), Tangipahoa(D, Westfall,1974), Washington (JL). Vidrine (1992) sight records from Beauregard (V), Rapides (V), Sabine (V), and Vernon should be questioned until voucher specimens are obtained. The Tangipahoa Parish specimen was collected by Sid Dunkle on 4 May 1986 at Kentwood, Tangipahoa River at Hwy 38. (May 4-Jun 6) [#E]

Gomphus (Gomphurus) vastus (Walsh, 1862). Cobra Clubtail. NEW STATE RECORD. Iberville: Mississippi River near Sr-75, Plaquemine Ferry Landing, 12,13,14 June-2005, collection and photos Gayle or Jeanell Strickland. Images: Male Female (Use <back> button to return to list )

Gomphus (Hylogomphus) apomyius Donnelly, 1966. Banner Clubtail. East Baton Rouge (B9,L), East Feliciana (B9). Tangipahoa (F,W), Washington (W). Since this species occurs in southeast Texas (Donnelly, 1966,1978; Orr, 1988), it probably will be collected in the pinelands of southwest Louisiana. ( Mar 25-Apr 24) [#U>] Images: Male Female (Use <back> button to return to list )

Gomphus (Gomphus) australis (Needham), 1897. Clearlake Clubtail. NEW STATE RECORD. St. Tammany: Snead Dr at SR-435, Hillcrest Pond, 15-Apr-2005, collection and photos Gayle or Jeanell Strickland. Images: Male (Use <back> button to return to list )

Gomphus (Gomphus) exilis Selys, 1854. (= Gomphus flavocaudatus Walker). Lancet Clubtail. Ascension (G), East Baton Rouge (F,JU,W), East Feliciana (JU,W), Livingston (F), St. Helena (F,W), St. Tammany ,Tangipahoa, Washington, West Feliciana (G). A very common early-season species in the southeast pinelands, it often flies with G. lividus. Although it has not been reported west of the Mississippi in Louisiana, Abbot (1996) reports it from Nacogdoches County, Texas, as a western extension of its range. ( Mar 3-May 24) [E] Images: Male Female (Use <back> button to return to list )

Gomphus (Gomphus) hodgesi Needham, 1950. Hodges' Clubtail. St. Tammany. So far it has been collected only in spring fed tributaries within the lower Pearl River drainage. Talisheek Creek at SR-41 is the best place to collect it.. This is the western limit of this lower coastal plain species, the range of which extends eastward into the Florida panhandle. ( Mar 1-Apr 3) [CP>w] ] Images: Male Female (Use <back> button to return to list )

Gomphus (Gomphus) lividus Selys, 1854. Ashy Clubtail. Beauregard (JL,L,V), Bienville (K5), East Feliciana (F,W), Evangeline (V), LaSalle (W), Lincoln, Natchitoches, Ouachita(M), Rapides (B,D,V), St. Helena (W), Tangipahoa, Vernon (D,V,V8,W), Washington (D,JL,J,W). Along with G. exilis, this is the most common early-season gomphid. ( Mar 13-Jun 6) [E] Images: Male Female (Use <back> button to return to list )

Gomphus (Gomphus) militaris Hagen in Selys, 1858. Sulphur-tipped Clubtail. Caddo: Reported for the first time by Ellzey (2004). (May 18) [W#]

Gomphus (Gomphus) oklahomensis Pritchard, 1935. Oklahoma Clubtail. Allen (V,W), Beauregard (F,VP), Bienville (K9), Caddo (J), Calcasieu (D,V), LaSalle (W), Lincoln (L), Natchitoches (K4), Rapides (B8,D), Vernon (D,V,V8,W), Winn (JA). G. oklahomensis replaces G. exilis west of the Mississippi River in Louisiana, where it is commonly found with G. lividus. ( Mar 23-May 2) [#U<e] Images: Male Female (Use <back> button to return to list )

Hagenius brevistylus Selys, 1854. Dragonhunter. Allen, Avoyelles (JU), Beauregard (JL,JU), Bossier (JU), East Baton Rouge (JU), East Feliciana, Grant, Jackson (JL), Lincoln (JU), Natchitoches (J), Rapides, St. Helena, St. Landry, St. Tammany, Tangipahoa, Vernon, Washington, West Feliciana (JU), Winn (JU). (May 10-Aug 7) [E] Images: Male Female (Use <back> button to return to list )

Ophiogomphus australis Car1e, 1992. Southern Snaketail. Tangipahoa, Washington (types, Carle, l992). This may be the only species that comes close to being a Louisiana endemic, as it has been found only in two parishes in southeast Louisiana and adjacent Pike County, Mississippi of the Tangipahoa River headwaters. It requires a gravel substrate. Gravel mining in southeast Louisiana has all but destroyed the habitat for this species. ( Apr 8-Apr 21) [#>] Images: Male Female (Use <back> button to return to list )

Progomphus obscurus (Rambur, 1842). Common Sanddragon. Allen, Beauregard (JL,V), Bossier, Caddo (L,JU), Catahoula (J,JU), Claiborne, East Baton Rouge, East Feliciana, Grant, LaSalle (JL,M,W), Livingston, Natchitoches (JL,JU,L), Rapides, Red River (J), Sabine (V), St. Helena (F,J), St. Tammany, Tensas (JU,VP), Tangipahoa, Union, Vernon, Washington, Webster (L), West Feliciana. This common late-spring to early-summer species should be found along almost any sandy flowing stream in the pinelands throughout the state. ( Apr 11-Aug 25) [E] Images: Male Female (Use <back> button to return to list )

Stylurus amnicola (Walsh, 1862). Elusive Clubtail. Washington, Livingston (JU). The Washington Parish record of one female was verified by L.K. Gloyd and M. J Westfall Jr. (Bick, pers. comm.). I did not see the Barr specimen from Livingston Parish. (June 2) [E>]

Stylurus laurae Williamson, 1932. Laura's Clubtail. Beauregard (V), Grant (SS), Jackson (JL), Rapides (S9), Vernon (V), Washington (JL).( Apr 6-Jul 28) [#E] Images: Male (Use <back> button to return to list )

Stylurus plagiatus (Selys, 1854). Russet-tipped Clubtail. Allen (V), Caldwell , Catahoula, East Baton Rouge (L), Evangeline (V), Lincoln (JU), Ouachita (M), St. Landry (V), St. Tammany, Washington (G), West Carroll. (Jun 14-Oct 19) [T] Images: Male Female (Use <back> button to return to list )

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Cordulegastridae

Cordulegaster bilineata (Carle, 1983). (= Zoraena bilineata). Brown Spiketail. Washington Parish, Varnado, SR-21 at Pushepatapa Creek, 9-April-1987, 1 male, J. Daigle; So. Franklinton, Hwy 25, ward 1, Sec 23 or 24, 25-March-1984, 1 male, C.B. Barr, det WFM. First records for Louisiana. [*CP>w] Images: Male Female (Use <back> button to return to list )

Cordulegaster erronea Hagen in Selys, 1878.(= Kalyptogaster erronea). Tiger Spiketail. West Feliciana (JL) This single record was based on a larva collected at "Tunica Bayou headwater trib. La Hwy 66, 21-April-1973." Louton (1982) shows its primary range as mostly northern, with the nearest southern records being near the south Tennessee border. Tennessen (1995) reported it from northeast Alabama, and Harp (pers. Comm.) reports it from Clark County Arkansas. [#E>w]

Cordulegaster maculata Selys,1854. (= Pangaeagaster maculata). Twin-spotted Spiketail . Bossier (VP), East Feliciana (JL), Jackson (J,JL), Natchitoches (JL), Rapides (D,F), St. Helena (F,L,W), St. Tammany, Webster, West Feliciana (JL). Louton (1982) discussed the deviation of Louisiana specimens from the ones of the eastern part of its range. He suggested that Louisiana, Texas, and Arkansas specimens might be different enough to warrant subspecies status and designated these as "ssp. A" ( Mar 2-Apr 14) [E] Images: Male (Use <back> button to return to list )

Cordulegaster obliqua fasciata Rambur, 1842. Arrowhead Spiketail. East Baton Rouge (JU), East Feliciana (JU), Lincoln [as C. fasciatus] (JU) Natchitoches (K4, K5), Webster [as C. fasciatus], Rapides (V), Vernon (V,V8). I did not see any of the above specimens. Since Harp & Harp (1996) report it from adjacent Miller and Columbia Counties, Arkansas; and Donnelly (1978) reports Cordulegaster obliqua obliqua from southeast Texas, C. obliqua fasciata is retained on the state list. Voucher specimens from Louisiana are needed. (Apr 8-May 20) [E] Images: Male Female (Use <back> button to return to list )

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Corduliidae -Macromiinae

Didymops transversa (Say, 1839). Stream Cruiser. Allen (W), Beauregard (V), Caddo (O), East Feliciana, Evangeline (V), Grant, Jackson (JU), LaSalle (M,W), Livingston (F), Natchitoches, Ouachita (M), Rapides, St. Helena (F,W), St. Tammany, Tangipahoa, Vernon (D,V), Washington, West Felciana (G), Winn (JU). ( Mar 08-Apr 17) [E] Images: Male Female (Use <back> button to return to list )

Macromia illinoiensis georgina (Selys, 1878). Georgia River Cruiser. (Nomenclature follows Donnelly & Tennessen (1994). Acadia (V), Allen (WW), Avoyelles (V), Beauregard (JL,V), Bossier (J), East Baton Rouge(Garcia, 1934,JU,W), East Feliciana (W), Evangeline (V), LaSalle, Livingston(D,JU), Ouachita (M,V as illinoiensis), Rapides (V), St. Helena (W), St. Landry (V), St. Tammany, Vernon (JA), Washington (D,JL), West Feliciana (W). ( May 12-Aug 31) [E] Images: Male Female (Use <back> button to return to list

Macromia taeniolata Rambur, 1842. Royal River Cruiser. Acadia (V), Beauregard (V), Caddo (JU), Catahoula, DeSoto (JU), East Baton Rouge (JL,JU), East Feliciana (JU), Jackson (JU), Lafayette, Livingston (JU), Madison, Morehouse, Natchitoches (JU), Ouachita, Rapides (L,JU), Richland (U), St. Helena (F), St. Landry (V), St. Tammany, Tangipahoa, Union, West Feliciana (JU,W), Winn (JL,JU), ( May 7-Aug 31) [E] Images: Male (Use <back> button to return to list

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Corduliidae- Corduliinae

Epitheca costalis (Selys, 1871). (= Tetragoneuria costalis ). Stripe-winged Baskettail. Calcasieu(D), St. Tammany(D), Tangipahoa (Tennessen, 1977), Washington (J). Because it is reported from Arkansas (Harp & Harp, 1996), it should also occur in northern Louisiana. (Mar-25-May 2 ) [#E] Images: Female (Use <back> button to return to list

Epitheca cynosura (Say, 1839). (= Tetragoneura cynosura ). Common Baskettail. Acadia (V), Allen (V,W), Beauregard (B,V), Caddo (B,J,JU,L), Calcasieu (V), Catahoula (J), DeSoto, East Baton Rouge (L,JU), East Feliciana (W), Evangeline (B,V), Iberia (B,V), Jackson, Jefferson, Jefferson Davis (V), Lafayette (B,V), LaSalle (J,W), Lincoln, Livingston (W), Natchitoches (B,JU, K4, L), Orleans, Ouachita(M), Pointe Coupee, Rapides, Red River (JU,L), St. Helena (W), St. John the Baptist, St. Landry (V), St. Martin, St. Tammany, Tangipahoa, Tensas (VP), Union (VP), Vernon (D,V,V8,W). Washington, Webster, West Feliciana, Winn.( Feb 17-May 8) [E] Images: Male Female (Use <back> button to return to list

Epitheca princeps (Hagen, 1861). (= Epicordulia princeps ). Prince Baskettail. Acadia (V), Allen, Assumption (L), Avoyelles, Beauregard (V), Bossier (J,JU), Caddo (J,JU), Calcasieu (V), Caldwell (JU), Catahoula, Claiborne, DeSoto (JU), East Baton Rouge (W), Evangeline (L,V), Iberville (JU,L), Jefferson, Jefferson Davis (V), Lincoln (L), Livingston (F,JU), Madison, Morehouse (L), Natchitoches, Ouachita, Rapides, St. Helena (W), St. Landry, St. Tammany (F), Tangipahoa (JU,VP), Tensas (JU,L,VP) Union, Webster (JU), West Baton Rouge (W), West Feliciana (J)( May 2-Aug 19) [E] Images: Male Female (Use <back> button to return to list )

Epitheca spinosa (Hagen, in Selys, 1878). (= Tetragoneuria spinosa). Robust Baskettail. Dunkle reports (per comm.) a single female in his collection with the following data: Washington Parish: Varnado, Pushepatapa Creek at Hwy 21, 9-April-1987. Tennessen (1994) reports it from southwest Arkansas and southeast Oklahoma, so it probably occurs in northern Louisiana. [*CP>]

Epitheca stella Williamson in Muttkowski, 1911. (=Tetragoneuria stella). Florida Baskettail. Muttokwski (1911) reported that Ris (pers. comm) had examined a specimen from Louisiana in the de Selys collection. Davis (1933) and Needham & Westfall(1955) listed it for the state. Mulhern (1971) reported it: Ouachita: US Hwy. 80, 11 ml E. of Monroe, 14-April-1969 (R.G. Kirkpatrick) 1 male, Specimen in Northeast Louisiana University collection, Monroe La. Present also in Adams County, Mississippi (Kormondy, 1960). [#CP]

Helocordulia selysii (Hagen in Selys, 1878). Selys' Sundragon. East Feliciana (F), Lincoln, Natchitoches (JL), Rapides, St. Helena, St. Tammany, Tangipahoa (F), Vernon (V) Washington (F). ( Mar 12-Apr 10) [CP] Images: Male (Use <back> button to return to list

Neurocordulia alabamensis Hodges in Needham & Westfall, 1955. Alabama Shadowdragon. Jackson (JL), St. Tammany, Tangipahoa, Washington. All these records are based on larvae, one of which was reared (Louton, 1982). [CPw]

Neurocordulia molesta (Walsh, 1863). Smoky Shadowdragon. Avoyelles (V), Bossier (L), East Baton Rouge (L), Evangeline (V), St. Helena (F), St. John the Baptist (J), St. Landry, Vernon (JL), Washington. ( May 20-Jul 9 ) [E]

Neurocordulia obsoleta (Say, 1839). Umber Shadowdragon. Bick (1957) summarized historical Louisiana records and speculated that the New Orleans record of Burmeister(1839) was in error. I found a specimen in the Barr Collection: Beauregard, Bayou Anacoco, 1-July-1978,1 male, collector unknown. This is the only verified specimen from the state. Although not a new record, this species is restored to the state list. It should be found in northwest Louisiana also, since there is a Columbia Co., Arkansas record (Harp & Rickett, 1977). [*E]

Somatochlora filosa (Hagen, 186l ). Fine-lined Emerald. Allen (V), East Baton Rouge (Garcia, 1934), St. Tammany (B9,D,L). Vernon (J). Originally reported by Barr (1981) as a state record with no specific data, along with the specimen described below under S. margarita. Harp (pers. comm.) reports it from Bradley County, Arkansas, so it may occur in north Louisiana. ( Jul 7-Sep 24) [#E]

Somatochlora georgiana Walker, 1925. Coppery Emerald. Vernon (Daigle,1994): Bayou Anacoco River, West of Pinewoods, 6-Aug-1954, 1 female. coll S. M Russel, in George H Beatty collection. The writer collected it in Jackson County, Mississippi. Both Daigle (1994) and Abbott (1996) reported it from Titus County, Texas, so it probably occurs elsewhere in the pinelands. [#CP]

Somatochlora linearis (Hagen, 1861). Mocha Emerald. Allen (V), Beauregard (V), Bossier (JU), Caddo (J), Caldwell, DeSoto (JU), East Baton Rouge (JU,L), East Feliciana (F,W), Jackson, Lincoln, Livingston (JU,L), Rapides (V), Red River (L), Sabine (J), St. Helena (F), St. Tammany, Union (J,L). Vernon (J,V), Washington (G), West Caroll (J), Winn. ( Jun 12-Aug-17) [E]

Somatochlora margarita Donnelly, 1962. Texas Emerald. Bienville: Redfield Branch, 16-June-1980. coll J.E.B, (As "provocans" by JEB). Barr reported S. provocans as a new state record with no specific data. The writer examined this. specimen and determined it to be margarita. Dunkle reports (pers. comm) the following data on a specimen he examined at the USNM : Jackson Parish: Hwy 556 near the Lincoln Parish line, 23-July-1973, 1 male. The only other state from which it has been reported is Texas (Donnelly, 1962). [*U<e]

Somatochlora provocans Calvert, 1903. Treetop Emerald. Dunkle reports (per comm.) a specimen from Washington Parish in USNM: Crain's Creek, 8-June 1974, coll. Louton, det SWD 1991. He indicated that this is a new early date for the species and the westernmost record for the species. [*E>w]

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Libellulidae

Brachymesia gravida (Calvert, 1890). (= Cannacria gravida) Four-spotted Pennant. Acadia (V), Allen, Ascension (L), Assumption (L), Beauregard (V), Caddo (L), Calcasieu (U,V), Cameron, East Baton Rouge, East Feliciana (G4), Evangeline (V), Iberia (B,V), Jefferson, Jefferson Davis (J,V), Lafayette, Lafourche, Natchitoches, Ouachita, Orleans, Plaquemines, Pointe Coupee, Rapides (L), Sabine (V), St. Bernard, St. Charles, St. James (F), St. John the Baptist (D,F,L), St. Landry (V), St. Martin (B,L), St. Mary (L,V), St. Tammany, Tangipahoa (L), Tensas, Terrebonne, Vermilion, Vernon (V,V8), West Feliciana (W). Often abundant in brackish marshes and ponds of the deltaic plain, less frequent northward in the state. ( Apr 27-Oct 24). [E] Images: Male Female (Use <back> button to return to list )

Celithemis amanda (Hagen, 1861). Amanda's Pennant. Beauregard (B), East Feliciana (W), Rapides (SS), St. Tammany, Vernon (V,V8,V8B) ( May 30-Sep 9). Collected in adjacent Union County, Arkansas (Harp & Harp, 1996), so probably occurs in north Louisiana also. [CP] Images: Male Female (Use <back> button to return to list )

Celithemis bertha Williamson, 1922. Red-veined Pennant. Allen (V), St. Tammany: Abita Springs, 4.2 ml NE, at U-V Light, May-Sept, 1984-86, V.Brou, 23m, 2 f (in FSCA). Vidrine (1988b) first reported it as a new state record with no specific data from the "Bogs and Baygalls of Central Vernon Parish" . He compared the habitat to those "described for similar ones in Southeast St.Tammany Parish" (Jul 5-Sep 23) [#CP] Images: Young Male Male (Use <back> button to return to list )

Celithemis elisa (Hagen, 1861). Calico Pennant. Allen (V), Beauregard (G,L,V), Caddo (L,JU), Calcasieu (V), Claiborne, East Baton Rouge (L,JU), East Feliciana (G), Jefferson Davis (V), Lincoln, Livingston (F,L,J,JU), Natchitoches (K4), Ouachita, Rapides (V), St. Helena (F,L), St. Tammany, Tangipahoa (F), Vernon (V), Washington. ( Mar 9-Sep 15) [E] Images: Young Male Male Female (Use <back> button to return to list )

Celithemis eponina (Drury, 1773). Halloween Pennant. Acadia (D,L,JU), Allen, Ascension (F,J,L), Avoyelles (L,J,JU), Beauregard (J), Caddo (O), Calcasieu (F,U,V), Cameron (B,V), Claiborne, East Baton Rouge, East Feliciana (W), Evangeline (V), Iberia (B), Jefferson, Jefferson Davis (D,J,JU), Lafayette, Lafourche (JU), LaSalle (W), Lincoln (J,JU,L), Morehouse, Natchitoches, Orleans, Ouachita, Plaquemines, Rapides (J,JU,L), Red River (J,JU,L), Sabine (VP), St. Bernard (W), St. John the Baptist (F,L), St. Landry (V), St. Martin (B), St. Mary (J,JU,L), St. Tammany, Tangipahoa (L,JU), Tensas, Terrebonne (J), Vermilion, Vernon (V,V8), Webster, West Feliciana (W) ( Apr 21-Sep 7) [E] Images: Male Female (Use <back> button to return to list )

Celithemis fasciata Kirby, 1889. Banded Pennant. Acadia (V), Allen (V), Beauregard (V), Caldwell (J), Claiborne, East Baton Rouge, East Feliciana (F), Evangeline (L), Grant, LaSalle (W), Lincoln (L,JU), Livingston (L,JU), Natchitoches (K4), Orleans, Ouachita, Rapides (V), Sabine (VP), St. Helena (F,W), St. Landry (V), St. Tammany, Tangipahoa (F), Vernon, Washington. (May 17-Sep 10) [E] Images: Male Male (form monomelaena) Female (Use <back> button to return to list )

Celithemis ornata (Rambur, 1842). Faded Pennant. Allen (V), Beauregard (B), Caddo (JU), St. Landry (JU), St. Tammany, Vernon (V,V8), Washington (W). ( Mar 12-Sep 9) [CP] Images: Male Female (Use <back> button to return to list )

Celithemis verna Pritchard, 1935. Double-ringed Pennant. Allen (JL,V) East Feliciana (F), Rapides (V), St. Tammany (D,F), Vernon (JL,V,V8,V8b), Washington (L,W). ( Apr 24-Jun 12 ) [#E] Images: Male (Use <back> button to return to list )

Dythemis velox Hagen, 1861. Swift Setwing. LaSalle (W), Natchitoches (B8), Rapides (S9), St. Helena (W). This species has expanded its range considerably in the last 30 years. When Bick and I did extensive collecting during the 50's and 60's, we did not see it. It is now fairly common in north Louisiana, especially at abandoned gravel pits. In southeast Louisiana it is now often found at roadside borrow pits and ponds, mainly to the east and north of Baton Rouge. ( Aug 11-Sep 8). [#E] Images: Male Female (Use <back> button to return to list )

Erythemis simplicicollis (Say, 1839). Eastern Pondhawk. Now reported from all 64 parishes. All but the following were listed in Bick (1957): Assumption (L,JU), Concordia (L,JU), East Carroll (L,JU), East Feliciana (L,JU), Red River (L,JU), Richland (L,JU), St. John the Baptist (F,JU,W), Union (L,JU). Common throughout the state. ( Feb 18-Nov 18) [E] Images: Male Female (Use <back> button to return to list )

Erythemis vesiculosa (Fabricius, 1775). Great Pondhawk. NEW STATE RECORD. Cameron: Johnston’s Bayou, 2 miles east nr SR-82, 1 male (G7). This widespread species is found throughout Mexico, Central and South America, as well as, the Antilles. Images: Male (Use <back> button to return to list )

Erythrodiplax berenice (Drury, 1770). Seaside Dragonlet. Calcasieu (V,V9), Cameron, East Baton Rouge (JU), Iberia (V,V9), Jefferson, Lafourche, Orleans, Plaquemines, St. Bernard, St. Mary (V,V9), St. Tammany, Tangipahoa (JU), Terrebonne, St. John the Baptist (W), Vermilion (V,V9). The East Baton Rouge specimen is probably a vagrant, as this is a true brackish water species. I found adults several miles inland from the marshes in the Slidell (St. Tammany Parish) area during periods of windy conditions blowing from the marshes. At Lake Catherine (Orleans Parish) and North Shore (St. Tammany Parish), adults were sometimes seen by the thousands on warm summer days. ( Apr 11- Nov 4) [CP] Images: Male Female banded wing form (Use <back> button to return to list )

Erythrodiplax minuscula (Rambur, 1842). Little Blue Dragonlet. Acadia (V), Allen, Assumption, Beauregard (B), Bienville, Calcasieu, Cameron (V), Claiborne, East Baton Rouge, East Feliciana (F,W), Evangeline (B,JU), Jefferson Davis, LaSalle (W), Lincoln, Livingston (F,J,JU), Natchitoches, Orleans, Ouachita, Rapides, St. Helena (F,J,JU,W), St. Landry (L,JU), St. Martin (B), St. Tammany, Terrebonne, Vermilion (L,V), Vernon (J,JU), Washington, Webster (L,JU), West Baton Rouge (J,JU,L) ( Apr 3-Dec 9) [E] Images: Male Young Male Female (Use <back> button to return to list )

Erythrodiplax umbrata (Linnaeus, 1758). Band-winged Dragonlet. Cameron (G7), Jefferson, Lafayette (F), Rapides (S9), St. Tammany. This species which is abundant throughout subtropical and tropical America, is rare in Louisiana. ( Jun 13- Oct 27) [E] Images: Male Female (Use <back> button to return to list )

Ladona deplanata (Rambur, 1842). (= Libellula deplanata). Blue Corporal. Acadia (V), Allen (V), Beauregard (V), Caddo (L,JU), Calcasieu (L,V), Caldwell (M), DeSoto (B), East Baton Rouge (F,JU), East Feliciana (F), Evangeline (B,V), Jefferson Davis (V), LaSalle (W), Lincoln, Natchitoches (K4), Rapides, St. Helena (F,W), St. Tammany (Bennefield, 1965,W),Union, Vernon (F,V,V8), Washington (D,J). ( Feb 21-May 6) [E] Images: Male Female (Use <back> button to return to list )

Libellula auripennis Burmeister, 1839. Golden-winged Skimmer. Acadia, Allen (L,J,V), Ascension (JU), Beauregard (J), Bossier (JU), Caddo (JU), Calcasieu (J,V), Cameron (JU), Claiborne, DeSoto (JU), East Baton Rouge (J,JU), East Feliciana (W), Evangeline (V), Iberia (V), Jefferson Davis (D,V), Livingston, Natchitoches (J), Rapides (V), St. Helena (F), St. John the Baptist (W), St. Landry (V), St. Mary (J,V), St. Tammany, Tangipahoa (B,JU), Vermilion (V), Vernon, Washington (J), Webster (JU). All historical records such as that of Wright (1943) from Orleans, are excluded because they were before Westfall (1943) differentiated needhami and auripennis. Except for one specimen reported by Bick (1957), I do not know of any valid records from the deltaic areas between New Orleans and New Iberia where he collected between 1950 and 1975. The single specimen from Bick (1957) was probably a vagrant from the pinelands north of Lake Pontchatrain where auripennis is common. Since auripennis and needhami look alike in the field, the Vidrine sight records need verification. (May 20-Sep 9) [E] ] Images: Male Female (Use <back> button to return to list )

Libellula axilena Westwood, 1837. Bar-winged Skimmer. Beauregard(D), St. Tammany, Washington. Bick (1957) doubted the validity of the Wright(1939) record from Orleans Parish. The Beauregard record is the westernmost verified record for the species. There are no verified specimens collected between the Washington-St.Tammany area and Beauregard in the west. Records of Barr(1979) [ Ascension, East Baton Rouge, Point Coupee, and Webster] are questionable, as the only specimens correctly labeled axilena by Barr in the LSU collection were from St. Tammany Parish. The rest were incesta or vibrans. (May 23-Sep 9) [Ew] ] Images: Male (Use <back> button to return to list )

Libellula cyanea Fabricius, 1775. Spangled Skimmer. Catahoula (J), Grant (SS), Jackson (J), Madison (Foster, 1915), Natchitoches (G4), Ouachita(M), Rapides (V), Union (J). Washington(D). The Foster (1915) record was a sight record. Bick included it in his "records of species not seen by the writer" category and commented that it probably does occur in northern Louisiana. I examined the Barr specimens from Livingston Parish listed in his "handout" (1979). The specimens were L. flavida. The surprising record is that of Dunkle in Washington Parish at Pushepatapa Creek at Hwy. 21, 3-May-1986, 1 male. This record is the southernmost for this common widespread species. It should be found at more localities in the northern half of the state. It is often found at farm ponds and beaver ponds. (May 3-Jul 28) [#E^] Images: (Use <back> button to return to list ) Images: Male Female (Use <back> button to return to list )

Libellula flavida Rambur, 1842. Yellow-sided Skimmer. Acadia (V), Allen (V), Beauregard (V), Calcasieu (V), Catahoula (J), East Baton Rouge (JU), East Feliciana, Evangeline (V), Jackson (J), Jefferson Davis (V), Lincoln, Pointe Coupee (JU), Rapides (D,V), Sabine, St. Landry (V), St. Tammany (L), Tangipahoa, Union, Vernon (D,V,V8), Washington (W), Webster (JU). This widespread species is very localized and never abundant in Louisiana. (Apr 24-Sep 6) [E] Images: young Male Male Female (Use <back> button to return to list )

Libellula incesta Hagen, 1861. Slaty Skimmer. Acadia(D,V), Allen, Ascension, Avoyelles, Beauregard, Bienville (J), Bossier, Caddo, Calcasieu, Caldwell, Catahoula, Claiborne, Concordia (L,JU), DeSoto (L,JU), East Baton Rouge (F,L,JU,W), East Feliciana, Evangeline, Grant, Iberia (V), Jackson, Jefferson Davis (D,V), Lafayette, Lafourche (J), LaSalle, Lincoln, Livingston (F), Madison, Morehouse, Natchitoches, Ouachita, Pointe Coupee (L,J,JU), Rapides, Red River (F,J), Sabine, St. Helena, St. Landry (J,L,V), St. Martin (J,V), St. Mary (L), St. Tammany, Tangipahoa (D,J,L,JU), Union, Vernon, Washington, Webster, West Carroll, West Feliciana, Winn. This very common and widespread species is absent from New Orleans and the surrounding deltaic plain. (May 09-Sep 25) [E] Images: Young Male Mature Male Female (Use <back> button to return to list )

Libellula luctuosa Burmeister, 1839. Widow Skimmer. Bossier, Caddo (K4, O), East Feliciana (G), Evangeline (V), Grant (SS), Lincoln (L), Ouachita, Rapides (V), Sabine (V), St. Landry (V), Tensas (L,JU), West Carroll (J), West Feliciana (G). Neither Bick (1957) nor I found verified speciemens in the southern half of the state. It has been reported in southeast Texas(Orr, 1988) but probably will not be found in southeast Louisiana. ( May 10-Aug 23) [E^] Images: Male Female (Use <back> button to return to list )

Libellula needhami Westfall, 1943. Needham's Skimmer. Acadia, Allen (L,JU), Ascension (F), Assumption, Beauregard (V), Bossier, Calcasieu, Cameron, East Baton Rouge, Evangeline (VP), Iberia, Jefferson, Jefferson Davis (V), Lafayette, Lafourche, Livingston (W), Orleans, Ouachita, Plaquemines, Pointe Coupe (L,J,JU), Rapides (VP), St. Bernard, St. Charles, St. James (F), St. John the Baptist (W), St. Landry (V), St. Martin (J), St. Mary, St. Tammany, Tangipahoa, Terrebonne, Vermilion, Vernon (V8). Bick (1957) discussed the distribution differences between auripennis and needhami, and reported huge concentrations of needhami in the New Orleans area. The writer noted the same when he lived there. ( Apr 24-Oct 18) [CP] Images: Male Female (Use <back> button to return to list )

Libellula pulchella Drury, 1773. Twelve-spotted Skimmer. Allen (JU), Caddo (O), Calcasieu(U), Catahoula, East Baton Rouge, East Feliciana (G), Lincoln, Livingston (L), Orleans, Ouachita, Pointe Coupee (JU), Richland(M), St. Tammany, Tangipahoa, Tensas(M), Washington, West Carroll. This widespread species is rare in Louisiana, at least in the southern half of the state. I did not see it in the New Orleans area and believe that the Ris (1910) and other historical records from the "New Orleans area" are actually from St. Tammany or Tangipahoa parishes just north of Lake Pontchartrain. The erroneous "previous records" of Barr (1979) and Vidrine (1992) are excluded. ( Mar 28-Nov 7) [T] Images: Female (Use <back> button to return to list )

Libellula semifasciata Burmeister, 1839. Painted Skimmer. Allen, Ascension (JU), Beauregard (J,V), Bossier (JU), Caddo (JU), Calcasieu (V), Claiborne, East Baton Rouge (L,JU), Evangeline (V), LaSalle (W), Lincoln, Livingston (F), Natchitoches (K4), Rapides (V), St. Tammany, Tangipahoa (F), Vernon (V,V8,W), Washington. ( Mar 6-Aug 25) [E] Images: Male Female (Use <back> button to return to list )

Libellula vibrans Fabricius, 1793. Great Blue Skimmer. Acadia (D,J,L), Allen, Ascension (F,J,JU,L), Assumption (J), Avoyelles, Beauregard (J,JU,L), Bienville (J,JU,L), Bossier, Caddo (JU,L), Calcasieu, Caldwell (J), Cameron (V), Catahoula, Claiborne (J,JU), Concordia, DeSoto (J,JU,L), East Baton Rouge, East Feliciana, Evangeline, Franklin (J),Grant, Iberia, Iberville (JU,L,W), Jefferson, Jefferson Davis, Lafayette, Lafourche, LaSalle, Lincoln, Livingston, Madison, Morehouse, Natchitoches (B,J,JU,L,), Orleans, Ouachita, Plaquemines, Pointe Coupee (JU,L), Rapides, Red River (J,JU,L), Richland (J), Sabine, St. Bernard, St. Charles, St. John the Baptist, St. Helena, St. James (JU,W), St. Landry, St. Martin (J), St. Mary, St. Tammany, Tangipahoa, Terrebonne, Union (J), Vermilion (V), Vernon, Washington, Webster (J,L), West Baton Rouge (JU,L), West Feliciana (J,L,W), Winn (B,L). ( Mar 26-Sep 25) [E] Images: Young Male Male Female (Use <back> button to return to list )

Macrodiplax balteata (Hagen, 1861). Marl Pennant. Acadia (V), Calcasieu (V), Cameron (V), Jefferson Davis (V) Lafourche (J), St. Helena (G), St. Tammany. Neither Bick (1957) nor I collected or saw this species in the field in Louisiana. Bick (1957) commented that he examined the 2 specimens collected at the old Lacombe fish hatchery. I examined a student- collected specimen from Fountainbleau State Park in 1964. Both of these St. Tammany Parish localities are only a few miles apart on the edge of the pinelands adjacent to brackish areas along the north shore of Lake Pontchatrain. Records of this species from other states indicates that it is associated with brackish water. Voucher specimens are needed from other parishes. Orr (1988) reported it from southeast Texas. ( May 10-Nov 18) [T] Images: Male Female (Use <back> button to return to list )

Miathyria marcella (Selys in Sagra, 1857). Hyacinth Glider. Acadia (V), Allen (V), Ascension (F), Beauregard (V), Calcasieu (V), East Baton Rouge, Iberville (F), Lafayette (B), Livingston (G), Natchitoches (K4), Orleans, Ouachita (Anna Hill), Pointe Coupee (L), Rapides (SS), St. Bernard (F), St. Helena (F), St. Landry (L,V), St. Martin (B,V), St. Tammany, Tangipahoa (J), Vermilion (V), West Feliciana. Most collections of this species have been in October and November. It seems to be restricted to the southern half of the state. It is usually found wherever water hyacinth (Eichhornia crassipes ) is growing, as it is its preferred breeding habitat (Bick, 1950) (Apr 13-Nov 24) [CP] Images: Male Female (Use <back> button to return to list )

Orthemis ferruginea (Fabricius, 1775). Roseate Skimmer. Acadia (V), Allen, Avoyelles (V), Beauregard, Calcasieu (V), Cameron (B,V), Claiborne, Concordia, DeSoto, East Baton Rouge, Evangeline (V), Iberville (L), Jackson (JU), Jefferson, Jefferson Davis (V), Lafayette, Livingston (F), Natchitoches (O), Orleans, Rapides (V), St. Landry (V), St. Tammany, Tangipahoa (G), Vermilion (V), Vernon, Washington (D), West Carroll. This widespread species was not very abundant anywhere. It appears to have two emergence peaks within Louisiana, one in the spring and one in the late summer-early fall. ( Apr 27-Dec 1) [T] Images: Male Female (Use <back> button to return to list )

Pachydiplax longipennis (Burmeister, 1839). Blue Dasher. Now reported from all 64 parishes. New records added since Bick (1957): Acadia (D,J,JU,L), Bienville (J,JU,L), Caldwell (J,JU,L), Concordia (J,JU,L), East Carroll (J,JU,L), Morehouse (J,JU,L), Pointe Coupee (J,JU,L), Red River (J,JU,L), St. James (JU,L). Bick (1957) stated that this was the most frequent and abundant odonate in Louisiana. ( Mar 9-Dec 2) [T] Images: Male Female (Use <back> button to return to list )

Pantala flavescens (Fabricius, 1798). Wandering Glider. Present in all parishes but Washington and West Baton Rouge. New records added since Bick (1957): Acadia (J,JU,L), Ascension (JU,W), Bienville (J,JU,L), Caddo (JU), DeSoto (JU,L), Evangeline (JU,L), Iberville (F,JU), Livingston (F,JU), Red River (J,JU), Richland (J,JU), Sabine (JU), St. Bernard (F,JU), St. Helena (JU), St. John the Baptist (F,JU), Vernon (JU,V8), Washington (G4), Webster (JU,L), West Baton Rouge (L). This common species is frequently encountered flying in parking lots and road intersections. On numerous occasions I encountered females ovipositing on the hood of my dusty dark blue car. (Apr 13-Dec 5) [T] Images: Male Female (Use <back> button to return to list )

Pantala hymenaea (Say, 1839). Spot-winged Glider. Allen (J), Beauregard (D,V), Bienville (JU,L), Bossier (JU,L), Caddo, Caldwell, Cameron, Claiborne (JU), DeSoto (JU), East Baton Rouge (JU,L), East Feliciana (W), Evangeline (V), Franklin (J), Grant, Jackson, Iberville (L), Jefferson Davis (JU,L), Lafayette, Lafourche, Lincoln (JU), Madison, Morehouse (JU,L), Natchitoches (O), Ouachita, Pointe Coupee, Rapides (V), Red River (J,JU), Richland (J), Sabine (V), St. John the Baptist (L), St. Landry (V), St. Martin (V), St. Tammany, Union, Vermilion (V), Vernon (V,V8), Webster (JU), West Feliciana (G4), Winn (J,JU). (Jun 6-Sep 9) [T] Images: Male Female (Use <back> button to return to list )

Perithemis tenera (Say, 1839). Eastern Amberwing. Recorded from all but 3 parishes: Cameron, Franklin, and Plaquemines. New records added since Bick (1957): Acadia (D,L,V), Bienville (L), Caddo (J,L), Iberia (B,V), Iberville (F,L), Jackson (L), Livingston (F,L), Richland (J), Sabine (V), St. Helena (L), St. John the Baptist (F), St. Landry (L), Terrebonne (L), West Baton Rouge (F,L). ( Feb 1-Nov 21) [E] Images: Male Female (Use <back> button to return to list )

Plathemis lydia (Drury, 1770). (= Libellula lydia). Common Whitetail. Acadia (V), Allen, Ascension (F,L,JU), Avoyelles (JU, L,), Beauregard, Bienville, Bossier, Caddo (J,JU,L,O), Calcasieu (J,V), Caldwell (J,JU), Catahoula, Claiborne, Concordia, DeSoto, East Baton Rouge, East Carroll, East Feliciana, Evangeline, Grant, Iberia (B), Iberville (F,L,JU), Jackson, Jefferson Davis (L,V), Lafayette, LaSalle, Lincoln, Livingston (F,L), Madison, Morehouse (J,L), Natchitoches, Orleans, Ouachita, Pointe Coupe (J,JU), Rapides, Red River, Richland, Sabine (L,V), St. Charles (F), St. Helena (F,L,W), St. James (J,JU), St. John the Baptist (F), St. Landry (L,V), St. Martin (B,V), St. Mary (L), St. Tammany, Tangipahoa, Tensas, Vermilion (V), Vernon (B,L,V,V8), Washington, Webster, West Baton Rouge (F,L,JU), West Carroll, West Feliciana, Winn (J,L). This is another species absent from the New Orleans area and most of the deltaic plain. The historical 19th century record from Orleans (Hagen, 1875) was probably from St. Tammany Parish. ( Mar 3-Nov 24) [E] Images: Young Male Male Female (Use <back> button to return to list )

Sympetrum ambiguum (Rambur, 1842). Blue-faced Meadowhawk. Caddo (K4), East Baton Rouge, Jackson (JU),Ouachita(M), Richland, St. Martin (B), St. Tammany, Winn (JU). Bick (1957) discussed the preference of this species for shaded swamps and sloughs. It should occur in southwest Louisiana also, as it is in southeast Texas (Orr, 1988). ( Jun 27-Nov 27) [E] Images: Male (Use <back> button to return to list )

Sympetrum corruptum (Hagen, 1861). (= Tarnetrum corruptum). Variegated Meadowhawk. Acadia (V), Allen (V), Ascension (W), Bienville (J), Caddo (J), Calcasieu (V), Cameron (V), East Baton Rouge, East Feliciana, Evangeline (V), Iberville (L), Jefferson, Jefferson Davis (V), Lafayette, Lincoln, Natchitoches, Orleans, Plaquemines, Rapides (B), Sabine (V), St. Bernard (F),. St. Charles, St. Landry (V), St. Tammany, Tangipahoa, Terrebonne, Washington (D). Wright(1943) stated that this species flew from April to late fall. Both Bick (1957) and the writer found this species only in the spring and the fall. Donnelly (1978) also reported two separate periods of adult activity in southeast Texas. (Mar 23-Apr 13 & Oct 31- Dec 20) [T] Images: Male Female (TX) (Use <back> button to return to list )

Tramea carolina (Linnaeus, 1763). Carolina Saddlebags. Allen, Ascension (JU,L), Beauregard, Caddo, Calcasieu (JA), Caldwell, Cameron, East Baton Rouge, East Feliciana (F), Evangeline (B), Grant, Iberville (G), Jackson, Jefferson (B), Jefferson Davis, Lafayette, Lincoln, Livingston, Madison, Morehouse (L), Natchitoches, Orleans, Plaquemines, Pointe Coupee (JU,L), Rapides (B,JU,L), St. Charles, St. Helena, St. John the Baptist, St. Tammany, Tangipahoa, Terrebonne, Vernon (V8), Washington, Webster (JU), West Feliciana (JU,L), Winn . ( Feb 24-Nov 15) [E] Images: Male Female (Use <back> button to return to list )

Tramea lacerata Hagen, 1861. Black Saddlebags. Acadia (V), Allen, Ascension (W), Avoyelles, Beauregard, Bossier (JU), Caddo (JU,O), Calcasieu, Caldwell (JU,L), Cameron, Claiborne, DeSoto (JU), East Baton Rouge (B,JU,L), Evangeline, Grant (JU), Iberia (V), Iberville (L), Jefferson, Jefferson Davis, Lafayette, Lafourche, Lincoln, Madison, Morehouse (JU), Natchitoches, Orleans, Ouachita, Plaquemines, Rapides, Red River (B,JU,L), Sabine (JU), St. Charles, St. John the Baptist(D,F), St. Landry (JU,L), St. Martin (V), St. Mary, St. Tammany, Tangipahoa (W), Tensas, Terrebonne, Union, Vermilion, Vernon (D,V,V8), Washington, Webster, West Feliciana. ( Mar 30-Nov 5) [T] Images: Male Female (Use <back> button to return to list )

Tramea onusta Hagen, 1861. Red Saddlebags. Acadia (J,L,V), Allen, Beauregard, Caddo, Calcasieu, Cameron, Claiborne (JU), East Baton Rouge (J,JU,L), Evangeline (V), Iberia, Jefferson Davis (V), Lafayette, Rapides (V), Madison, Morehouse (JU), Orleans (B), Ouachita (M), St. Landry (V), St. Martin (V), St. Mary, St. Tammany, Vermilion, Vernon (V,V8), West Feliciana (JU), Winn. ( Mar 31-Oct 30) [T] Images: Male Female (Use <back> button to return to list )

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Additions and corrections to list (since Jan 1998) Many. Parish records in Red

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Doubtful Records (Includes vagrants)

Louisiana records of the following are doubtful because of one or more of the following reasons: misidentification, synonymy, indefinite locality, no recent collections, vagrants, and/or out of known range. Included here are a few unpublished records from institutional collections (e.g. Amphiagrion saucium). They are included as a reference for future researchers, in case the records are cited in future publications. A few of these species are also placed in the "Expected Species" list below:

Lestes forcipatus Rambur, 1842. Reported from "lower Louisiana" (Foster & Smith, 1901; Foster, 1915 ). Louisiana is well south of the known range and Bick(1957) considered that Louisiana records should be L. australis,. Both Walker (1952) and Westfall & May (1996) discussed the taxonomic confusion between forcipatus and australis.

Lestes rectangularis Say, 1839. Reported from "lower Louisiana" by Foster & Smith (1901). There are no specimens of this species from the state, but it is expected in Louisiana (see below).

Lestes unguiculatus Hagen, 1861. Cameron (Hine, 1906). This locality is far out of the known range. This widespread northern transcontinental species has been reported from Arkansas and Oklahoma by Westfall & May (1996), but not from any Gulf Coast state. If it occurs in Louisiana, it should be only in the northern half of the state.

Lestes vidua (Hagen, 1861). Often listed for Louisiana because of the New Orleans type locality which is far west of the accepted range. I speculate that this specimen was mis-labeled. Westfall & May (1996) considered it doubtful for Louisiana and reported the nearest record being from Alabama.

Amphiagrion saucium (Burmeister, 1839). There are no published records of the species from the state. I include it here to in case the record is inadvertently published in the future. In NMNH there are 6 males and 2 females in the pinned specimen collection, labeled "N.O.L.A.//Shufeldt; RPC 1912" . Dr. Flint checked the determinations, so the specimens must be mis-labeled as to locatilty, as the species has not been recorded anywhere along the Gulf Coast, the nearest record being from north Alabama (Tennessen, 1995).

Argia barretti Calvert, 1902. Vidrine (1992) reported it from Vernon Parish based on a single specimen photographed at Fort Polk . After careful examination of the slide, it could not be verifed as barretti, a species that occurs in south Texas and Mexico.

Enallagma aspersum (Hagen. 1861). Reported from "lower Louisiana" by Foster & Smith (1901). There are no verified records of this species, but it is expected in Louisiana (see below).

Enallagma pollutum (Hagen 1861) Reported from "lower Louisiana" by Foster and Smith (1901). Bick (1957) pointed out that this was before Calvert's (1919) description of the similar species, E. vesperum. There are currently no verified records from Louisiana, but it is expected (see below).

Enallagma praevarum (Hagen, 1861) Bick (1957) discussed the history of this record. There are no verified records east of a line running north-south through San Antonio, Texas either Texas or Oklahoma.

Ischnura verticalis (Say, 1839). Madison (Foster, 1915). Reported for the state by Westfall & May (1966) and by Bick (1957) under "Species not seen by the Writer." There are no modern records for I. verticalis. It may occur in northern Louisiana, but all Arkansas records are from the northern third of Arkansas (Harp, pers. comm.); and Abbott (pers. comm.) has no adjacent records in Texas.

Nehalennia irene (Hagen, 1861). Reported from Madison Parish (Foster, 1915), probably without knowledge of Calvert's 1913 description of integricollis. The known range of N. irene is far north of Louisiana (Westfall & May, 1996).

Nehalennia pallidula Calvert, 1913. Reported from Grand Isle, Jefferson Parish (Behre, 1950). This species is known only from southern Florida in the Everglades region (Westfall & May, 1996). Bick (1957) reported that George Beatty had a female in his collection from Grand Isle. At my request, Clark Shiffer examined the Beatty specimen and verified its identity. The specimen is actually a male. The data label states "Grand Isle Research station", "Plaquemine La", "in rear porch". There is no habitat suitable for N. pallidula in either Plaquemine or southern Jefferson Parish. The specimen may have either been mislabeled or blown in by a hurricane.

Anax amazili (Burmeister, 1839). Orleans. Calvert (1927) reported two female specimens collected in 1918 in the city of New Orleans by Ris. In spite of abundant modern collecting in the New Orleans area, no one has collected A. amazili; therefore either the larva or the adult probably came in on a ship at the port of New Orleans. Dunkle (1989) described it as a "vagrant" to southern Florida, the Dry Tortugas, and Texas. Its normal range is Mexico and the West Indies, south to Argentina. Donnelly (Pers. Comm.) collected it in west Texas. Bick (pers. comm.) stated that the Louisiana record probably was based on a very rare stray. I consider this species, along with the next two, as vagrants or strays, not having an established population.

Coryphaeschna viriditas Calvert,1952. The only Louisiana record is a female at NMNH with the following data: "New Orleans, Tulane University, 5-11-58 , at light." Dunkle (1989) reported it from the southern tip of Florida and the Keys. Its range is listed from northern Mexico and the Greater Antilles south to Paraguay. The NMNH specimen was young and had perfect wings, so it was probably imported as an egg or larva and emerged in the Tulane University area of New Orleans.

Gynacantha hyalina Selys, 1882 .Williamson (1923) recorded a male in NMNH from Crowley LA. as very close if not identical to the widely distributed oriental G. hyalina. He stated that if it was really taken in Louisiana it was probably brought to the US as an egg or larva. This pinned specimen has the following data labels: 1st: "Crowley La.//7.17.11" 2nd: "ESTucker// ", 3rd: "4", 4th: "Cur//Gynacantha //hyalina// Selys" (across end): "examined//EBW 1920" [the word "hyalina" is scratched out by pencil], 5th: "very close// to hyalina E.B.W.10/6/20// locality label?"

Aphylla protracta (Hagen in Selys, 1859). Reported by Vidrine (1992), a determination assuredly based on Needham & Westfall (1955), but Garrison (1986) assigns this citation to A. angustifolia.

Arigomphus lentulus (Needham, 1902). Needham & Westfall (1955) listed it for Louisiana. Westfall (pers. comm, 1953,1957) considered that the record should probably refer to A. submedianus. Montgomery (1927) listed Gomphus sp."in the pallidus group" from Madison Parish, and Westfall (pers. comm., 1970) stated that the specimen may be A. maxwelli. Vidrine (1992) listed unacceptable sight records from 6 Parishes. Therefore there are no confirmed Louisiana records of lentulus. (See below under Expected Species)

Arigomphus paIIidus (Rambur, 1842). All Louisiana references to this species, discussed in detail in Bick (1957), were prior to the description of maxwelli and should refer to that species.

Gomphus (Gomphurus) fraternus (Say, 1839). Reported by Vidrine (1992) as a historical record. Louisiana is far southwest of the known range of fraternus.

Gomphus (Gomphurus) vastus Walsh, 1862. Vidrine (1992) gives unacceptable sight records for Allen and Beauregard parishes; nevertheless, it may occur in Louisiana (see below under expected species).

Gomphus (Gomphus) abbreviatus Hagen in Selys, 1878. Reported from "lower Louisiana" (Foster and Smith ,1901). This northeastern species reaches south only to South Carolina. The only Gomphus from this group found in Louisiana is apomyius which was not described until 1966 (Donnelly). It could have been this species.

Gomphus (Gomphus) flavocaudatus Walker, 1940. Synonym of G. exilis (see Garrison, 1997).

Gomphus (Gomphus) minutus Rambur, 1842. Originally reported from "lower Louisiana" (Foster & Smith, 1901) This is far west of the known range of this species.

Gomphus (Gomphus) spicatus Hagen in Selys, 1854. Reported from "lower Louisiana"(Foster & Smith, 1901). Louisiana is far south of the known range of this northeastern species.

Ophiogomphus mainensis Packard, 1863. Bick (1957) reported it from Tangipahoa and Washington Parishes. Carle (1992) stated that these records should be his O. australis.

Cordulegaster diastatops (Selys, 1854). (= Zoraena diastatops) Originally reported from the state by Walker (1958) before Carle's 1983 description of bilineata. Carle reported diastatops from several northeastern states and the southernmost record was from Virginia.

Helocordulia uhleri (Selys, 1871). Needham & Westfall (1955) and others have listed H. uhleri (Selys, 1871) from Louisiana; however, there are no specimens in FSCA, and all LA specimens of the genus examined were H. selysii. Bick(1957), in a footnote, discussed the confusion arising from the switched figures in Needham and Westfall (1955).

Neurocordulia virginiensis Davis, 1927. Bick(1957) reported one larva from St. Tammany Parish, but he now considers this as probably a misidentification (pers. comm.).

Epitheca (Tetragoneuria) semiaquea (Burmeister 1839). Reported by Vidrine (1992). While on a field trip with Vidrine, I collected several specimens that he identified as T. semiaquea. These specimens collected were T. cynosura. (see also under Expected Species)

Epitheca (Tetragoneuria) spinigera Selys, 1871. Bick(1957) discussed this record, and he concluded that it was too far out of its range. Dunkle (pers. comm.) states that this record was based on a misdetermination.

Libellula exusta Say, 1839. Reported from "lower Louisiana" (Foster & Smith, 1901). Bick (1957) stated that Louisiana records of exusta should be deplanata, as exusta is a northern species.

Nannothemis bella (Uhler, 1857). Reported from "lower Lousiana" (Foster & Smith, 1901). This is another species with no verified record, but it may occur in Louisiana (see also under Expected Species).

Perithemis seminole Calvert 1907. Reported from Orleans Parish ( Kormondy, 1960). This is a synonym of tenera.

Sympetrum vicinum (Hagen, 1861). Reported by Carle (1993) from Louisiana. Carle (pers. comm.) cannot recall the source of the data. (see also under Expected Species)

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Expected Species:

This category is presented here to assists future researchers with locating new records. These species have a reasonable probability of occurring in Louisiana, based on verified records from either Mississippi, Arkansas, or East Texas, within 100 miles of the Louisiana border. Some of the species on this list are also included in the doubtful records list, due to lack of a verified records.

Lestes rectangularis (Say, 1839). Bick (pers. comm.) reports it from adjacent Hancock County, MS, so it may occur in the Washington-St.Tammany Parish area.

Enallagma aspersum (Hagen, 1861). (See also "Doubtful Records"). Present in nearby Lawrence County, MS (F). U.S. distribution suggests that it may occur in northern Louisiana or even in the Washington Parish area.

Enallagma doubledayi (Selys, 1850). Recorded from adjacent Marion County, MS (Lago et al, 1980), just north of Washington Parish. (Now recorded from Louisiana 2004, see list above)

Enallagma pallidum (Root, 1923). Present in adjacent Hancock County, MS (F). It may occur in the St. Tammany-Washington Parish area..

Enallagma pollutum (Hagen, 1861). (See also "Doubtful Records"). There is a specimen from adjacent Hancock County, MS, in FSCA, so it possibly occurs in the St. Tammany -Washington Parish area.

Arigomphus lentulus (Needham, 1902). (See also "Doubtful Records). There is a record of lentulus from Hunt County, TX (Harwell, 1951), and it is also reported from southeast Texas (Donnelly, 1978; Orr, 1988), so it could turn up in western Louisiana.

Gomphus (Gomphurus) vastus Walsh, 1862. (See also "Doubtful Records"). Louisiana is entirely within the accepted range of this widespread eastern species. Nearby records include Rankin County, MS (F) and Franklin County, AR (F). and widespread in Texas (Donnelly, pers. comm.) (Now recorded from Louisiana 2005, see list above)

Gomphus( Gomphus) militarus Hagen in Selys, 1858. Reported from Marion County, TX (Harwell, 1951), and southeast Texas (Donnelly,1978). It may occur in northwest and extreme west central Louisiana. (Now recorded from Louisiana 2004. See list above)

Stylurus potulentus (Needham, 1942). Reported from Pearl River County, MS (Bick, 1983). It could occur in St. Tammany -Washington Parish area.

Stylurus townesi Gloyd, 1936. Pearl River County, MS (Bick, 1983). It could occur in the St. Tammany-Washington Parish area.

Epitheca (Tetragoneuria) semiaquea (Burmeister 1839). Donnelly (pers. comm.) reports this species as "common" in east Texas. Look for it in Louisiana.

Neurocordulia xanthosoma (Williamson, 1908). Harwell (1951) recorded it from adjacent Marion County, TX. It may occur in northwest Louisiana.

Nannothemis bella (Uhler, 1857). (See also "Doubtful Records"). Present nearby in George County, MS (Westfall, l952), and Jackson County, MS (F), which is about 75 miles east of the Pearl River/Louisiana border.

Sympetrum vicinum (Hagen, 1861 ). (see also "Doubtful Records). Collected in nearby Bradley County, AR (Harp, pers. comm.) but only from west and central Texas (Dunkle, pers. comm.). It may occur in north Louisiana.

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Bibliography

Includes references from Louisiana, as well as, Arkansas, Mississippi, and east Texas, and general references.

Abbott, J. C. 1996. New and interesting records from Texas and Oklahoma. Argia 8(4): 14-15

Adams, C. C. 1900. Odonata from Arkansas. Ent. News. 11(10): 621-622

Banks, N. 1892. A synopsis, catalogue, and bibliography of the neuropteroid insects in temperate North America. Trans. Amer. Ent. Soc. 19: 327-373

Barr, J. E. 1979. List of distribution records of Odonata (Anisoptera) in Louisiana. Unpub. personal list distributed at the annual meeting of the Louisiana Academy of Sciences.

Barr, J. E. 1981. Two new state records for Odonata of Louisiana. Proc. La. Acad. Sci. 44: 164 (abstract)

Behre, E. H. 1950. Annotated list of the fauna of the Grand Isle Region. Occ. Pap. Mar. Lab. L. S. U. 6: 1-65

Bell, R. and W. H. Whitcomb. 1961. Erythemis simplicicollis (Say) A dragonfly predator of the bollworm moth. Fla. Ent. 44(2): 95-97

Bell, R. and W. H. Whitcomb. 1966. Additional records of the Anisopterous Odonata in Arkansas. Unpub. manuscript

Bennefield, B. L. 1965. A taxonomic study of the genus Ladona (Odonata: Libellulidae). Univ. Kan. Sci. Bull. 45: 361-396

Bick, G. H. 1941. Life-history of the dragonfly Erythemis simplicicollis (Say). Ann. Ent. Soc. Amer. 34: 215-230

Bick, G. H. 1949. Night collections of dragonflies. Ent. News. 60: 182

Bick, G. H. 1950. The dragonflies of Mississippi (Odonata: Anisoptera). Amer. Mid. Natur. 43: 66-78

Bick, G. H. 1951a. The early nymphal stages of Tramea lacerata Hagen (Odonata: Libellulidae). Ent. News. 62: 293

Bick, G. H. 1951c. The nymph of Libellula semifasciata Burmeister. Proc. Ent. Soc. Wash. 53: 247-250

Bick, G. H. 1953b. The occurrence of Anax longipes in Mississippi. Ent. News. 64: 230-232

Bick, G. H. 1957. The Odonata of Louisiana. Tulane Stud. Zool. 5: 71-135

Bick, G. H. 1959. Additional dragonflies (Odonata) from Arkansas. Southwestern Nat. 4(3): 131-133

Bick, G. H. 1978. New state records of United States Odonata. Notul. Odonatol. 1: 17-36

Bick, G. H. 1990. Unpublished records in Florida State Collection of Arthropods (FSCA). Argia(1-4): 1-2

Bick, G. H. and J. F. Aycock. 1950. The life history of Aphylla williamsoni Gloyd. Proc. Ent. Soc. Wash. 52: 26-32

Bick, G. H., J. F. Aycock and A. Orestano. 1950. Tauriphilia australis (Hagen) and Miathyria marcella (Selys) from Florida and Louisiana. Proc. Ent. Soc. Wash. 52: 81-84

Bick, G. H. and J. C. Bick. 1957. The Odonata of Oklahoma. Southwestern Nat. 2: 1-18

Bick, G. H., L. E . Hornuff and E. N. Lambremont. 1953. An ecological reconnaissance of a naturally acid stream in southern Louisiana. Jour. Tenn. Acad. Sci. 28: 221-231

Borror, D. J. 1942. A revision of the Libelluline genus Erythrodiplax (Odonata). Ohio St. Univ. Grad. Sch. Stud. Contr. Zool & Ent. 4: xx+286

Bridges, C. A. 1991. Catalogue of the family-group, genus-group, and species-group names of the Odonata of the World. Charles A. Bridges, Urbana, IL. L+708pp

Byers, C. F. 1937. A review of the dragonflies of the genera Neurocordulia and Platycordulia. Misc. Pub. Mus. Zool. Univ. Michigan 36: 1-35

Byers, C. F. 1939b. A study of the dragonflies of the genus Progomphus (Gomphoides) with a description of a new species. Proc. Fla. Acad. Sci. 4: 19-85

Carle, F. L.1979a. Two new Gomphus (Odontata:Gomphidae) from eastern North America with adult keys to the subgenus Hylogomphus. Ann. Ent. Soc. Am. 72(3): 418-426

Carle, F. L.1993a Ophiogomphus (Ophionurus) australis spec. nov. from the Gulf Coast of Louisiana, with larval and adult keys to American Ophiogomphus (Anisoptera: Gomphidae). Odonatologica. 21(2): 141-152

Carle, F. L. 1993b. Sympetrum janeae spec. nov. from eastern North America, with a key to the Neartic Sympetrum (Anisopter: Libellulidae). Odonatologica 22(1): 1-16

Carle, F. L. 1983. A new Zoraena (Odonata:Cordulegaster) from eastern North America, with a key to the adult Cordulegaster of America. Ann. Ent. Soc. Am.76(1): 61-68

Cook, Carl (198?) Report on a study of Macromia dragonflies held by the USNM. Unpublished USFW report. Projects #31420-1115 and 31440-1115

Daigle, J. J. 1994. The larva and adult male of Somatochlora georgiana Walker (Odonata: Corduliidae). Bull. Amer. Odon. 2(2): 21-26.

Davis, W. T. 1933. Dragonflies of the genus Tetragoneuria. Bull. Brooklyn. Ent. Soc. 28(3): 87-104

Delcarpio, J. B., R. B. Baerwald and L. J. Magnuson, 1983 Multiple SR-T tubule junctions in a single insect flight muscle fiber. J. Ultrastr. Res. 84(2): 151-160

Donnelly, T.W. 1962. Somatochlora margarita, A new species of dragonfly from eastern Texas (Odonata: Corduliidae). Proc. Ent. Soc. Wash. 64(4): 235-240

Donnelly, T. W. 1964. Enallagma westfalli, A new damselfly from eastern Texas, with remarks on the genus Teleallagma Kennedy. Proc. Ent. Soc. Wash. 66(2): 103-109

Donnelly, T. W. 1966. A new Gomphine dragonfly from eastern Texas. (Odonata: Gomphidae). Proc. Ent. Soc. Wash. 68(2): 102-105

Donnelly, T. W. 1973. The status of Enallagma traviatum and westfalli (Odonata: Coenagrionidae). Proc. Ent. Soc. Wash 75: 297-302

Donnelly, T. W. 1978. Odonata of the Sam Houston National Forest and vicinity, east Texas, United States, 1960-1966. Notul.Odonatol.1(1): 6-7

Donnelly, T. W. 1992. The Odonata of New York. Bull. Amer. Odonatol. 1(1):1-27

Dunkle, S. W. 1975. New records of North American anisopterous dragonflies. Fla. Ent. 58: 117-119

Dunkle, S. W. 1983. New records of North American Odonata. Ent. News. 94(4): 136-138

Dunkle, S. W. 1989. Dragonflies of Florida Peninsula, Bermuda, and the Bahamas. Scientific Publishers, Gainesville Fl. 154 pp

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